Urocaridella renatekhalafae Khalaf, 2018: New Species of Cave Clear Cleaner Shrimp from Gulf of Oman by Dr-Norman Khalaf

Urocaridella renatekhalafae Khalaf, 2018 : A New Cave Clear Cleaner Shrimp Species from the East Coast of the United Arab Emirates, Gulf of Oman, Arabian Sea, Indian Ocean ‫ نوع جديد من روبيان الكهوف‬: 8102 ، ‫أوروكاريدّال ريناته خلفي خلف‬ ، ‫ال ُمنظف الشفاف من الساحل الشرقي لدولة اإلمارات العربية ال ُمتحدة‬ ‫ ال ُمحيط الهندي‬، ‫ بحر العرب‬، ‫خليج عُمان‬ By : Prof. Dr. Sc. Norman Ali Bassam Ali Taher Mohammad Ahmad Mostafa Khalaf-Prinz Sakerfalke von Jaffa & Ola Mostafa Esmail Mostafa Khalaf (Photography)

‫ فً بحر‬8 ‫) من مولع السفٌنة الغارلة إنشكٌب‬8102 ، ‫روبٌان رٌناته خلف ال ُمنظف الشفاف (أوروكارٌدّال رٌناته خلفً خلف‬ 0100108101 .. ‫ عدسة ُمدربة الغواصٌن وأستاذة التصوٌر التحت مائً الكابتن عُال خلف‬.. ‫ مترا‬88 ‫خورفكان على عُمك‬ The New Species : Renate Khalaf‟s Cave Clear Cleaner Shrimp (Urocaridella renatekhalafae Khalaf, 2018) from the ship wreck Inchcape 2 in Khorfakkan Sea at a depth of 22 meters .. Photo by PADI Dive Instructor & Underwater Photography Instructor Ola Khalaf .. 14.10.2017 .. https://jaffacity.wixsite.com/gazellebulletin/urocaridella-renatekhalafae-khalaf--1 Gazelle : The Palestinian Biological Bulletin – Number 157 – January 2018

Abstract: A new species of the Cave Clear Cleaner Shrimp genus Urocaridella (Crustacea: Decapoda: Palaemonidae) from the East Coast of the United Arab Emirates, Gulf of Oman, Arabian Sea, Indian Ocean is described. This species is distinguished from the other six Urocaridella species by its very long rostrum and their distinctive transparent body covered by numerous striking small white and red dots and rings. It occurs in shallow sub-tidal waters to a depth of 24 meters. It is morphologically and geographically distinct from the other Clear Cleaner Shrimp species. The new species was named Urocaridella renatekhalafae Khalaf, 2018.

Keywords: Arthropoda, Crustacea, Malacostraca, Decapoda, Palaemonidae, Urocaridella renatekhalafae, New Species, Cave Clear Cleaner Shrimp, Renate Khalaf’s Cave Clear Cleaner Shrimp, Renate Khalaf, Khorfakkan, Sea of Khorfakkan, Emirate of Sharjah, Fujairah, Sea of Fujairah, Emirate of Fujairah, United Arab Emirates, Gulf of Oman, Arabian Sea, Indian Ocean, Near East, Middle East.

‫ فً بحر‬8 ‫) من مولع السفٌنة الغارلة إنشكٌب‬8102 ، ‫روبٌان رٌناته خلف ال ُمنظف الشفاف (أوروكارٌدّال رٌناته خلفً خلف‬ .0100108101.. ‫ عدسة ُمدربة الغواصٌن وأستاذة التصوٌر التحت مائً الكابتن عُال خلف‬.. ‫ مترا‬88 ‫خورفكان على عُمك‬

The New Species : Renate Khalaf‟s Cave Clear Cleaner Shrimp (Urocaridella renatekhalafae Khalaf, 2018) from the ship wreck Inchcape 2 in Khorfakkan Sea at a depth of 22 meters .. Photo by PADI Dive Instructor & Underwater Photography Instructor Ola Khalaf .. 14.10.2017 .. https://www.facebook.com/photo.php?fbid=1977941599125882&set=pb.100007301827082.2207520000.1515335752.&type=3&theater Gazelle : The Palestinian Biological Bulletin – Number 157 – January 2018

Introduction: On Friday 05.05.2017 & Saturday 14.10.2017, my wife the PADI Scuba Dive Open Water Instructor and Underwater Photography Instructor Ola Mostafa Khalaf managed to shoot magnificent photos of 3 cm. long undescribed Cave Clear Cleaner Shrimps from the Inchcape 10 ship wreck Sea Ground, 24 meters deep in the Sea of Fujairah, and the Inchcape 2 ship wreck Sea Ground, 22 meters deep in the Sea of Khorfakkan, East Coast of the United Arab Emirates, Gulf of Oman, Arabian Sea, Indian Ocean. These photos triggered our search for its identification. We examined zoological reference books and the internet searching for its identity. This new creature was not scientifically described and was not named. The closest genus for our new species was Urocaridella. Six species were studied and compared: Urocaridella antonbruunii Bruce, 1967, Urocaridella cyrtorhyncha Fujino & Miyake, 1969, Urocaridella liui Wang, Chan & Sha, 2015, Urocaridella pulchella Yokes & Galil, 2006, Urocaridella urocaridella Holthuis, 1950 & Urocaridella vestigialis Chace & Bruce, 1993. After studying and comparing it was identified as the undescribed Urocaridella sp. 1. Urocaridella is a small genus of shrimps with currently six species although many researchers claim that there are many more left to be discovered and described. Some of these tiny mosquito-like shrimps clean the mouths and bodies of moray eels and various other cave dwelling fish that share its habitat. Unlike Lysmata and Periclemenes, Urocaridella are capable of prolonged hovering and free swimming, and prefer doing so than perching (Tyk, 2013; Steinke, 2015). One of the neatest things about Urocaridella is their comical way of swimming. When swimming or hovering, they hold their claws downward and buzz around in an almost mosquito-meets-hovercraft fashion (Tyk, 2013; Steinke, 2015). They commonly swim by vigorously beating their pleopods when moving towards their fish clients. This behaviour gives them the strange appearance of hovering in mid-water (Calado, 2008). Sometimes they are called the Helicopter Shrimps.

Description and Distinctive Features Urocaridella renatekhalafae Khalaf, 2018 reach a length of 3.00 cm. The large adult female is almost transparent with white and red dots on the cephalothorax and abdomen. The legs have red and white rings and bands. The first and second pair of legs (pereopods) has white and red small claws, the first pair being smaller than the second pair. Small individuals are less pigmented than large ones, and the overall colour is probably less noticeable at night than during the day. The internal organs like the cephalothorax, stomach, digestive gland and ovary are often observed due to transparency; the ovary has an orange or light green color depending on the development of vitellogenesis. The cephalothorax has a very long white coloured rostrum bent upwards and largely exceeds the antennal scales. The eyes are normally stalked. The antennas and antennules are long and red, with some white dots at the base. Gazelle : The Palestinian Biological Bulletin â&#x20AC;&#x201C; Number 157 â&#x20AC;&#x201C; January 2018

Pereopods (walking legs) are long and slender. The pleopods (swimming feet under the abdomen) are effective and allow the animal to remain immobile in open water. When swimming, only the pleopods are in action, the pereopods hang under the animal. The abdomen forms a characteristic right angle at the level of which is a red spot preceded by a white spot. The red spot may widen to form a band that runs laterally through the abdomen. At the base of the telson, there is red and white bands; followed by red and white dots and rings on the uropod.

Identification Key Transparent body with white and red dots on the cephalothorax and abdomen. Slender, long legs with white and red rings and bands. Long red antennas and antennules. Very long white rostrum. Telson and uropod with white and red dots and rings.

Gazelle : The Palestinian Biological Bulletin – Number 157 – January 2018

Size Urocaridella renatekhalafae grows to about 3.00 cm long.

Diet Urocaridella renatekhalafae feeds on parasites that live on fish and moray eels, copepods, food waste and mouth phlegm. It attracts its host by a lateral vibratory movement of its red antennas and antennules. When the fish is set, it lands on the bottom near the shrimp. Then, the shrimp swims towards the fish and begins the cleaning. It sometimes enters the fish‟s mouth and gill cavities.

Habitat It is mainly observed from the surface up to 25 m depth. Renate Khalaf's shrimp loves rocky environments with holes for hiding.

Biology Transparent colouration in Urocaridella renatekhalafae is seen as a first line defence against predation. In its natural environment, the Renate Khalaf‟s Cave Clear Cleaner Shrimp lives in groups in small caves and crevices on the reef in locations called "cleaning stations”. It attracts its host by a lateral vibratory movement of its red antennas and antennules. When the fish is set, it lands on the bottom near the shrimp. Then, the shrimp swims towards the fish and begins the cleaning. It sometimes enters the fish‟s mouth and gill cavities. These shrimps are regularly observed in the company of the moray eels and fish which they clean. They can change host easily, including a diver, while swimming in open water. The biology of this unusual shrimp does not appear to have been researched by scientists. This shrimp forms groups of a few individuals in small caves or in the immediate vicinity; they have their cleaning station there and constitute a “guild” (solidarity association).

Reproduction The eggs are fixed on the pleopods. As this shrimp occurs in warm waters in its distribution range, it is likely to reproduce all year round.

Distribution The species Urocaridella renatekhalafae was described and scientifically named from the Fujairah and Khorfakkan Sea, East Coast of the United Arab Emirates, Gulf of Oman, Arabian Sea. Its distribution extends from the Red Sea, through the Indian Ocean and the Western Pacific Ocean. Gazelle : The Palestinian Biological Bulletin – Number 157 – January 2018

ً‫ ف‬01 ‫) من مولع السفٌنة الغارلة إنشكٌب‬8102 ، ‫روبٌان رٌناته خلف ال ُمنظف الشفاف (أوروكارٌدّال رٌناته خلفً خلف‬ 1.01.08101 .. ‫ عدسة ُمدربة الغواصٌن وأستاذة التصوٌر التحت مائً الكابتن عُال خلف‬.. ‫ مترا‬81 ‫بحر الفجٌرة على عُمك‬

The New Species : Renate Khalaf‟s Cave Clear Cleaner Shrimp (Urocaridella renatekhalafae Khalaf, 2018) from the ship wreck Inchcape 10 in Fujairah Sea at a depth of 24 meters .. Photo by PADI Dive Instructor & Underwater Photography Instructor Ola Khalaf .. 05.05.2017 ..

Classification Kingdom: Animalia Linnaeus, 1758 Phylum: Arthropoda Lankester, 1904 Class: Malacostraca Latreille, 1802 Order: Decapoda Latreille, 1802 Family: Palaemonidae Rafinesque, 1815 Subfamily: Palaemoninae Rafinesque, 1815 Genus: Urocaridella Borradaile, 1915 Species: Urocaridella renatekhalafae Khalaf, 2018

Etymology / Derivation of the Scientific Name The genus name Urocaridella derives from Greek Oura meaning tail, and Latin Carid meaning marine crustacean; and the species name renatekhalafae is Latin for “Renate Khalaf” (Mother of the Taxonomist, born in 1941). Gazelle : The Palestinian Biological Bulletin – Number 157 – January 2018

This new Cave Clear Cleaner Shrimp species was classified and named by the Palestinian-German Zoologist Prof. Dr. Sc. Norman Ali Bassam Khalaf-von Jaffa in January 2018, and the specific name „renatekhalafae‟ was named in honor of his beloved mother “Renate Margarete Khalaf geb. Weber” (born in Saarbrücken, Germany in 1941).

The New Species : Renate Khalaf‟s Cave Clear Cleaner Shrimp (Urocaridella renatekhalafae Khalaf, 2018) from the ship wreck Inchcape 10 in Fujairah Sea at a depth of 24 meters ... Photo by PADI Dive Instructor & Underwater Photography Instructor Ola Khalaf ... 05.05.2017. https://www.facebook.com/photo.php?fbid=1910797192506990&set=a.1408184479434933.1073 741829.100007301827082&type=3&theater

Conclusion: After examining the new Cave Clear Cleaner Shrimp species with white and red dots and rings, living off the East Coast of the United Arab Emirates, and referring to many zoological references, and searching the Internet, I came finally to a conclusion that we are in front of a new Clear Cleaner Shrimp species from the Gulf of Oman, Arabian Sea, Indian Ocean. Gazelle : The Palestinian Biological Bulletin – Number 157 – January 2018

I gave it the scientific name Urocaridella renatekhalafae, new species. The species name “renatekhalafae” is Latin for my mother “Renate Khalaf” (born in Saarbrücken, Germany in 1941). Urocaridella renatekhalafae, new species: Scientific binomial name: Urocaridella renatekhalafae Khalaf, 2018. Authority: Prof. Dr. Sc. Norman Ali Bassam Khalaf-von Jaffa. Common Names: Renate Khalaf’s Cleaner Shrimp, Renate Khalaf’s Cave Clear Cleaner Shrimp. Holotype: Urk-1, 3.00 cm., Prof. Dr. Norman Ali Khalaf-von Jaffa’s Collection. Location: Sea of Fujairah, Emirate of Fujairah, United Arab Emirates. Depth: 24 meters. Date of capture: 05th May, 2017.

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Hankyu Communications, Tokyo, 173 pp. Khalaf, Norman Ali Bassam (1980). Tabie‟t Al-Talawon fi Al-Haywanat (The Colouration of Animals). Al-Biology Bulletin. Number 1. January 1980, Safar 1401. Biological Society, Kuwait University, State of Kuwait. pp. 4-5. (In Arabic). Khalaf, Norman (Translator) (1982). Al-Miah Al-Mulawatha Tohaded Al-Asmak Bi‟ilinqiraad (Water Pollution threatens the Fish Fauna with Extinction). Al-Biology Bulletin. Number 18, Third Year, First Semester, Saturday 6.11.1982. Biological Society, Kuwait University, State of Kuwait. pp. 7. (Translation from German into Arabic). Khalaf, N.A.B. (1986). The Schooling of Sumatra Barbs (Barbus tetrazona tetrazona) and Minnows (Phoxinus phoxinus). Dissertation, Master of Science in Ecology, Departments of Zoology and Botany, University of Durham, England. September 1986. pps. 59 + iv. http://etheses.dur.ac.uk/7084/ Khalaf-von Jaffa, Norman Ali Bassam (1992). An Introduction to the Animal Life in Palestine. Gazelle. Gazelle: The Palestinian Biological Bulletin. Bonn-Bad Godesberg, Federal Republic of Germany. Number 30, Tenth Year, October 1992. pp. 1-7. (In Arabic). Khalaf-von Jaffa, Norman Ali Bassam (1994). An Introduction to the Animal Life in Palestine. Shqae‟q Al-Nouma‟n (Anemone coronaria). A Quarterly Magazine Issued by the Program EAI (Education for Awareness and for Involvement). Environmental Education / Children for Nature Protection. In Cooperation with Dept. of General and Higher Education. P.L.O., Palestine. Number 4. Huzairan (June) 1994. pp. 16-21. (In Arabic). Khalaf, Norman Ali Bassam (2001). The Extinct and Endangered Animals in Palestine. In: Gazelle: The Palestinian Biological Bulletin Home Page. Extinct and Endangered Animals and Reintroduction. http://gazelle.8m.net/photo3.html Khalaf-von Jaffa, Norman Ali (2004). Gazelle: Das Palästinensische Biologische Bulletin. Eine Wissenschaftliche Reise in Palästina, Arabien und Europa zwischen 1983 – 2004. / Gazelle: The Palestinian Biological Bulletin. A Scientific Journey in Palestine, Arabia and Europe between 1983 – 2004. ISBN 3-00-014121-9. Erste Auflage / First Edition, Juli 2004: 452 Seiten / Pages. Zweite erweiterte Auflage (Second Extended Edition), August 2004: 460 Seiten / Pages. Norman Ali Khalaf, Bonn-Bad Godesberg, Germany. http://dr-norman-ali-khalaf-books.webs.com/& eBook: https://www.yumpu.com/en/document/view/59405994/book-gazelle-thepalestinian-biological-bulletin-a-scientific-journey-in-palestine-arabia-and-europebetween-1983-2004-by-norman-ali-khalaf-von-jaffa-2004 Khalaf-von Jaffa, Norman Ali (2005). Aquatica Arabica. An Aquatic Scientific Journey in Palestine, Arabia and Europe between 1980 - 2005 / Aquatica Arabica. Eine Aquatische Wissenschaftliche Reise in Palästina, Arabien und Europa zwischen 1980 - 2005. ISBN 300-014835-3. Erste Auflage / First Edition, August 2005: 376 Seiten / Pages. Norman Ali Khalaf, Rilchingen-Hanweiler, Bundesrepublik Deutschland & Sharjah, United Arab Emirates. http://dr-norman-ali-khalaf-books.webs.com/aquaticaarabica.htm & eBook: https://www.yumpu.com/en/document/view/59407769/book-aquaticaarabica-an-aquatic-scientific-journey-in-palestine-arabia-and-europe-between-1980Gazelle : The Palestinian Biological Bulletin – Number 157 – January 2018

2005-by-norman-ali-khalaf-von-jaffa-2005 & eBook: https://joom.ag/nN6L Khalaf, N.A.B. (2005). The Schooling of Sumatra Barbs (Barbus tetrazona tetrazona) and Minnows (Phoxinus phoxinus). [M.Sc. Dissertation in Ecology, Departments of Zoology and Botany, University of Durham, England. September 1986. pps. 59 + iv]. In: Aquatica Arabica. An Aquatic Scientific Journey in Palestine, Arabia and Europe between 1980 2005. Erste Auflage, August 2005. Norman Ali Khalaf, Rilchingen-Hanweiler, Bundesrepublik Deutschland & Sharjah, United Arab Emirates. pp. 28-93. Khalaf-von Jaffa, Norman Ali Bassam (Member of PALESTA) (2005). Palestinian Scientists and Technologists Abroad (PALESTA). Gazelle: The Palestinian Biological Bulletin. Number 47, Twenty-third Year, November 2005, Shawal 1426. pp. 11-12. Sharjah, United Arab Emirates. (In Arabic). Khalaf, Norman Ali (2005, 2006, 2007). Chapter 3: Geography, Flora and Fauna. Pages 32-39. In: Palestine: A Guide. By Mariam Shahin, Photography by George Azar. CoAuthor: Norman Ali Khalaf. Northampton, Massachusetts: Interlink Publishing Group, 2005, 2006, 2007. xi + 471 pages. Appendices to page 500. http://ipsnewsite.mysite4now.com/journals.aspx?id=7323&jid=1&href=fulltext Khalaf-Sakerfalke von Jaffa, Norman Ali Bassam Ali Taher (Gründer) (seit Juni 2007). Yahoo! Deutschland Group: Fauna Palaestina. http://de.groups.yahoo.com/group/Fauna_Palaestina/ Khalaf-Sakerfalke von Jaffa, Norman Ali Bassam Ali Taher (2007). Felidae Arabica. A Zoological Journey in Palestine, Arabia and Europe between 1980-2007 / Felidae Arabica. Eine Zoologische Reise in Palästina, Arabien und Europa zwischen 1980-2007. ISBN 978-3-00-019568-6. Erste Auflage (First Edition), Juli (July) 2007, 300 pp. SelfPublisher: Norman Ali Khalaf, Rilchingen-Hanweiler, Deutschland & Sharjah, United Arab Emirates. (in Arabic, German and English). Doctoral Dissertation. Ashwood University, USA. Doctor of Science Degree in Zoology (Summa Cumm Laude) on 26.09.2007. http://dr-norman-ali-khalaf-books.webs.com/felidaearabica.htm & Doctoral Dissertation (eBook) : https://www.yumpu.com/en/document/view/59397999/doctorate-dissertationfelidae-arabica-by-norman-ali-bassam-khalaf-doctor-of-science-ashwood-universityusa-2007 Khalaf-Sakerfalke von Jaffa, Norman Ali Bassam Ali Taher (Gründer) (seit September 2007). Yahoo! Deutschland Group: Fauna Arabica. http://de.groups.yahoo.com/group/Fauna_Arabica/ Khalaf-Sakerfalke von Jaffa, Norman Ali Bassam Ali Taher (2007). Haywanat Filistin ‫( حٌوانات فلسطٌن‬Fauna of Palestine). Wikipedia, Al-Mawsu'a Al-Hurra (The Free Encyclopedia). Gazelle: The Palestinian Biological Bulletin. Number 69, Twenty-fifth Year, September 2007 CE, Sha‟ban 1428 AH. pp. 1-4. (in Arabic). http://ar.wikipedia.org/wiki/%D8%AD%D9%8A%D9%88%D8%A7%D9%86%D8%A7 %D8%AA_%D9%81%D9%84%D8%B3%D8%B7%D9%8A%D9%86 Khalaf-Sakerfalke von Jaffa, Dr. Sc. Norman Ali Bassam Ali Taher (2009). Flora and Fauna in Palestine. Gazelle: The Palestinian Biological Bulletin. ISSN 0178 – 6288. Number 91, July 2009, Rajab 1430 AH. pp. 1-31. Sharjah, United Arab Emirates. Gazelle : The Palestinian Biological Bulletin – Number 157 – January 2018

http://flora-fauna-palestine.webs.com/ Khalaf-von Jaffa, Dr. Norman Ali Bassam (2009). Fauna Palaestina – Part One. A Zoological Journey in Palestine, Arabia and Europe between 1983 – 2006 / Fauna Palaestina – Teil Eins. Eine Zoologische Reise in Palästina, Arabien und Europa zwischen 1983 – 2006. ISBN 978-9948-03-865-8. Erste Auflage/First Edition, September 2009: 412 Seiten/Pages. Self Publisher: Dr. Norman Ali Bassam Khalaf-von Jaffa, Sharjah, United Arab Emirates & Rilchingen-Hanweiler, Bundesrepublik Deutschland. http://dr-norman-ali-khalaf-books.webs.com/faunapalaestinapart1.htm & eBook: https://www.yumpu.com/xx/document/view/59498633/fauna-palaestina-1-bookby-dr-norman-ali-khalaf-2009 Dr. Sc. Norman Ali Bassam Ali Taher Khalaf-Sakerfalke von Jaffa, Zoologist, Ecologist and Geologist : The Scientific References (1980-2009). http://dr-norman-ali-khalafreferences.webs.com/ Khalaf-von Jaffa, Dr. Sc. Norman Ali Bassam Ali Taher (2010). Fauna Emiratus - Part One. Zoological Studies in the United Arab Emirates between 2004 - 2009. / Fauna Emiratus – Teil Eins. Zoologische Studien in die Vereinigten Arabischen Emirate zwischen 2004 - 2009. ISBN 978-9948-15-462-4. Erste Auflage/First Edition, November 2010: 350 Seiten / Pages. Self Publisher: Dr. Norman Ali Bassam Khalaf-von Jaffa, Dubai and Sharjah, United Arab Emirates & Rilchingen-Hanweiler, Bundesrepublik Deutschland. http://dr-norman-ali-khalaf-books.webs.com/faunaemiratuspart1.htm & eBook: & https://www.yumpu.com/xx/document/view/59546804/fauna-emiratus-part-1zoological-studies-in-the-united-arab-emirates-between-2004-2009-by-dr-norman-alibassam-khalaf-von-jaffa-2010 Khalaf-von Jaffa, Dr. Norman Ali Bassam (2012). Fauna Palaestina – Part Two. Zoological Studies in Palestine between 1983 – 2009 / Fauna Palaestina - Teil Zwei. Zoologische Studien in Palästina zwischen 1983 – 2009. ISBN 978-9948-16-667-2. 1. Auflage / First Edition : July 2012, Shaaban 1433 H. 208 Seiten / Pages (Arabic Part 120 Pages and the English Part 88 Pages). Publisher: Dar Al Jundi Publishing House, Jerusalem, Palestine. http://dr-norman-ali-khalafbooks.webs.com/faunapalaestinapart2.htm & eBook: https://www.yumpu.com/en/document/view/59602956/fauna-palaestinapart-2-book-by-dr-norman-ali-khalaf-von-jaffa-2012 Khalaf-von Jaffa, Dr. Norman Ali Bassam (2013). Fauna Palaestina – Part Three. Zoological Studies in Palestine between 2005 – 2012 / Fauna Palaestina - Teil Drei. Zoologische Studien in Palästina zwischen 2005 – 2012. ISBN 978-9950-383-35-7. Erste Auflage / First Edition : July 2013, Shaaban 1434 H. 364 pages (English Part 350 Pages and the Arabic Part 14 Pages). Publisher: Dar Al Jundi Publishing House, Jerusalem, State of Palestine. http://dr-norman-ali-khalafbooks.webs.com/faunapalaestinapart3.htm Khalaf-Sakerfalke von Jaffa, Prof. Dr. Sc. Norman Ali Bassam Ali Taher (2013). Garra rufa wadiqana Khalaf, 2013: A New Freshwater Doctor Fish Subspecies from Wadi Qana Nature Reserve, Salfit Governorate, State of Palestine. Gazelle: The Palestinian Gazelle : The Palestinian Biological Bulletin – Number 157 – January 2018

Biological Bulletin. ISSN 0178 – 6288. Number 103, July 2013, Ramadan 1434 AH. pp. 125. Dubai and Sharjah, United Arab Emirates. http://palestinefishes.webs.com/palestine-doctor-fish Khalaf-Sakerfalke von Jaffa, Prof. Dr. Sc. Norman Ali Bassam Ali Taher (2013). † Macropomoides palaestina Khalaf, 2013 : A New Coelacanth Fish Fossil Species from the Anthracothere Hill in Al-Naqab, Palestine. Gazelle: The Palestinian Biological Bulletin. ISSN 0178 – 6288. Number 107, November 2013, Muharram 1435 AH. pp. 30-38. Dubai and Sharjah, United Arab Emirates. http://quastenflosser.webs.com/macropomoidespalaestina.htm Khalaf-von Jaffa, Prof. Dr. Norman Ali Bassam (2014). Fauna Palaestina – Part Four. Zoological Studies in Palestine between 1983 – 2014 / Fauna Palaestina - Teil Vier. Zoologische Studien in Palästina zwischen 1983 – 2014. ISBN 978-9950-383-77-7. Erste Auflage / First Edition : July 2014, Ramadan 1435 H. 456 Pages (English Part 378 Pages and the Arabic Part 78 Pages). Publisher: Dar Al Jundi Publishing House, Al-Quds (Jerusalem), State of Palestine. http://fauna-palaestina-part1.webs.com/faunapalaestina4.htm Khalaf-von Jaffa, Prof. Dr. Norman Ali Bassam (2014). † Macropomoides palaestina Khalaf, 2013 : A New Coelacanth Fish Fossil Species from the Anthracothere Hill in Al-Naqab, Palestine. In: Fauna Palaestina – Part Four. Zoological Studies in Palestine between 1983 – 2014 / Fauna Palaestina - Teil Vier. Zoologische Studien in Palästina zwischen 1983 – 2014. ISBN 978-9950-383-77-7. Erste Auflage / First Edition : July 2014, Ramadan 1435 H. pp. 126 - 138. Publisher: Dar Al Jundi Publishing House, Al-Quds (Jerusalem), State of Palestine. Khalaf-Sakerfalke von Jaffa, Prof. Dr. Norman Ali Bassam Ali Taher (2015). Plants and Animals unique to Palestine. Gazelle: The Palestinian Biological Bulletin. ISSN 0178 – 6288. Number 125, May 2015. pp. 1-18. Dubai and Sharjah, United Arab Emirates. http://flora-fauna-palestine-2.webs.com/ Khalaf-von Jaffa, Prof. Dr. Norman Ali Bassam (2015). Fauna Palaestina – Part Five. Zoological Studies in Palestine between 1983 – 2016 / Fauna Palaestina - Teil Fünf. Zoologische Studien in Palästina zwischen 1983 – 2016. ISBN 978-9950-383-92-0. Erste Auflage / First Edition : July 2015, Ramadan 1436 H. 448 pp. (English Part 304 Pages and the Arabic Part 144 Pages). Publisher: Dar Al Jundi Publishing House, Al-Quds (Jerusalem), State of Palestine. http://fauna-palaestina-books.webs.com/ Khalaf-Sakerfalke von Jaffa, Prof. Dr. Sc. Norman Ali Bassam Ali Taher (2016). Haywanat Falastin (Fauna of Palestine) ‫ حٌوانات فلسطٌن‬. Gazelle: The Palestinian Biological Bulletin. ISSN 0178 – 6288. Number 144, December 2016, pp. 1-18. Dubai and Sharjah, United Arab Emirates (In Arabic). http://animals-of-palestine-2.webs.com/fauna-ofpalestine-arabic Khalaf-Sakerfalke von Jaffa, Prof. Dr. Sc. Norman Ali Bassam Ali Taher Mohammad Ahmad Mostafa & Ola Mostafa Esmail Mostafa Khalaf (2017). Diaphorodoris olakhalafi Khalaf, 2017 : A New Dorid Nudibranch Sea Slug Species from the Sea of Khorfakkan, Gulf of Oman, Arab Sea, Indian Ocean. Gazelle: The Palestinian Biological Bulletin. ISSN 0178 – 6288. Number 149, May 2017, pp. 31-47. Dubai and Sharjah, United Arab Gazelle : The Palestinian Biological Bulletin – Number 157 – January 2018

Emirates. http://marine-life-uae.webs.com/diaphorodoris-olakhalafi Khalaf-von Jaffa, Prof. Dr. Norman Ali Bassam (May 2017). Ola Khalaf‟s Nudibranch (Diaphorodoris olakhalafi Khalaf, 2017). EOL: Encyclopedia of Life. http://eol.org/collections/131000 Khalaf-von Jaffa, Prof. Dr. Norman Ali Bassam (May 2017). Species Diaphorodoris olakhalafi Khalaf, 2017. BioLib.cz. Biological Library Classification. http://www.biolib.cz/en/adtaxon/id406783/ Khalaf-Sakerfalke von Jaffa, Prof. Dr. Sc. Norman Ali Bassam Ali Taher Mohammad Ahmad Mostafa & Ola Mostafa Esmail Mostafa Khalaf (Photography) (2017). Trapania norakhalafae Khalaf, 2017: A New Dorid Nudibranch Sea Slug Species from the Dibba Sea, East Coast of the United Arab Emirates, Gulf of Oman, Arabian Sea, Indian Ocean. ‫ الساحل الشرلً لدولة اإلمارات‬، ‫ نوع جدٌد من بُزاق البحر عاري الخٌاشٌم من بحر دِبا‬: 2017 ، ‫ترابانٌا نورة خلفً خلف‬ ‫ ال ُمحٌط الهندي‬، ‫ بحر العرب‬، ‫ خلٌج عُمان‬، ‫ العربٌة ال ُمتحدة‬. Gazelle: The Palestinian Biological Bulletin. ISSN 0178-6288. Number 153, September 2017, pp. 7-30. Dubai and Sharjah, United Arab Emirates. (In Arabic & English). http://nudibranch-uae.webs.com/trapanianorakhalafae Khalaf-von Jaffa, Prof. Dr. Norman Ali Bassam (September 2017). Nora Khalaf‟s Nudibranch (Trapania norakhalafae Khalaf, 2017). EOL: Encyclopedia of Life. http://eol.org/collections/132727 Khalaf-von Jaffa, Prof. Dr. Sc. Norman Ali Bassam (September 2017). Taxon Profile: Species Dibba Brown-spotted Dorid Nudibranch Trapania norakhalafae Khalaf, 2017. BioLib.cz. Biological Library Classification. https://www.biolib.cz/en/adtaxon/id407709/ Khalaf-Sakerfalke von Jaffa, Prof. Dr. Sc. Norman Ali Bassam Ali Taher Mohammad Ahmad Mostafa & Ola Mostafa Esmail Mostafa Khalaf (2017). Presence of the Sexy Anemone Shrimp (Thor amboinensis de Man, 1888) in the Sea of Kalba, Gulf of Oman, Arab Sea, Indian Ocean. Gazelle: The Palestinian Biological Bulletin. ISSN 0178 – 6288. Number 156, December 2017, pp. 1-17. Dubai and Sharjah, United Arab Emirates. http://marine-life-uae.webs.com/sexy-shrimp Khalaf-Sakerfalke von Jaffa, Prof. Dr. Sc. Norman Ali Bassam Ali Taher Mohammad Ahmad Mostafa & Ola Mostafa Esmail Mostafa Khalaf (2018). Presence of the Banded Coral Shrimp (Stenopus hispidus Olivier, 1811) around the Snoopy Island, Emirate of Fujairah, United Arab Emirates, Gulf of Oman, Arab Sea. Gazelle: The Palestinian Biological Bulletin. ISSN 0178 – 6288. Number 157, January 2018, pp. 1-13. Dubai and Sharjah, United Arab Emirates. http://marine-life-uae-2.webs.com/ Khalaf-Prinz Sakerfalke von Jaffa, Prof. Dr. Sc. Norman Ali Bassam Ali Taher Mohammad Ahmad Mostafa & Ola Mostafa Esmail Mostafa Khalaf (2018). Urocaridella renatekhalafae Khalaf, 2018 : A New Cave Clear Cleaner Shrimp Species from the East Coast of the United Arab Emirates, Gulf of Oman, Arabian Sea, Indian Ocean. Gazelle: The Palestinian Biological Bulletin. ISSN 0178 – 6288. Number 157, January 2018, pp. 1431. Published by Prof. Dr. Norman Ali Khalaf Department for Environmental Research and Media, National Research Center, University of Palestine, Gaza, State of Palestine. https://jaffacity.wixsite.com/gazellebulletin/urocaridella-renatekhalafae-khalaf--1 Gazelle : The Palestinian Biological Bulletin – Number 157 – January 2018

Khalaf-Prinz Sakerfalke von Jaffa, Prof. Dr. Sc. Norman Ali Bassam (January 2018). Taxon Profile: Species Renate Khalaf‟s Cleaner Shrimp Urocaridella renatekhalafae Khalaf, 2018. BioLib.cz. Biological Library Classification. https://www.biolib.cz/en/adtaxon/id411136/

Khalaf, Ola (02.04.2017). Sexy Shrimp (Thor amboinensis). Facebook. https://www.facebook.com/photo.php?fbid=1880941872159189&set=a.14081844794349 33.1073741829.100007301827082&type=3&theater Khalaf, Ola (16.05.2017). Banded Coral Shrimp (Stenopus Hispidus). Facebook. Gazelle : The Palestinian Biological Bulletin – Number 157 – January 2018

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Gazelle : The Palestinian Biological Bulletin – Number 157 – January 2018