PG Biological Sciences by Edinburgh Global

Biological Sciences Postgraduate opportunities Prospectus 2013 entry www.ed.ac.uk

THE UNIVERSITY of EDINBURGH

Welcome to the University of Edinburgh: Influencing the world since 1583 Our proud history and alumni ambassadors For more than 400 years our people have been making their mark on the world. They’ve explored space, revolutionised surgery, won Nobel Prizes, published era-defining books, run the country, paved the way for life-saving breakthroughs and laid the foundations for solving the mysteries of the universe. By choosing further study or research at Edinburgh you will be joining a community of scholars who have been at the forefront of knowledge since 1583. We are associated with 15 Nobel Prize winners, including physicists Charles Barkla and Max Born, medical researcher Peter Doherty, economist Sir James Mirrlees and biologist Sir Paul Nurse. Our famous alumni include NASA astronaut Piers Sellers, former MI5 Director-General Dame Stella Rimington, Olympians Sir Chris Hoy and Katherine Grainger and historical greats such as philosopher David Hume, physicist and mathematician James Clerk Maxwell, inventor Alexander Graham Bell and Sherlock Holmes creator Sir Arthur Conan Doyle.

Teaching and research excellence We are consistently ranked as one of the world’s top 50* universities. As host to more than 30,000 students from some 130 countries, studying across 100 academic disciplines, the University of Edinburgh continues to attract the world’s greatest minds. World-leading research is produced by 96 per cent** of our academic departments, placing Edinburgh in the top five in the UK for research. Our excellent teaching was also confirmed in the latest report from the Quality Assurance Agency, which awarded us the highest rating possible for the quality of the student learning experience.

Collaborations and international partnerships As an internationally renowned centre of academic excellence, Edinburgh is the site of many world-class research collaborations. Our postgraduate students are crucial to our continued success and development and, along with our staff, they forge research links through regular travel and overseas exchanges. We take pride in our partnerships with other institutions such as the California Institute of Technology, Stanford University, the University of Melbourne, Peking University, the University of Delhi and the University of KwaZulu-Natal – to name but a few.

The University of Edinburgh Biological Sciences Postgraduate opportunities Prospectus 2013 entry

Linking research and commerce Edinburgh was one of the first UK universities to actively develop commercial links with industry, government and the professions. Edinburgh Research and Innovation (ERI) has continued, for the past four decades, to develop the promotion and commercialisation of the University’s research excellence. ERI assists our postgraduates in taking a first step to market, whether it is through collaborative research, licensing technology or providing consultancy services.

Enhancing your career With the best track record for graduate employment in the Russell Group, the University of Edinburgh is committed to embedding employability into the teaching and learning experience. From offering access to volunteering schemes to providing support from our sector-leading Careers Service, the University gives students myriad opportunities to develop the skills, knowledge and experience to give them the edge in a competitive job market.

An inspiring destination Your first-class education will take place in one of Europe’s most striking capital cities, which is regularly voted one of the best places in the world to live. Edinburgh enjoys a solid reputation as a centre for innovation, whether as home to the 18th-century Scottish Enlightenment or as a modern source of pioneering science, medicine and technology. You couldn’t ask for a more inspiring setting in which to further your knowledge and broaden your horizons.

Join us Edinburgh offers unparalleled academic breadth and diversity, making it a vibrant, challenging and stimulating environment for postgraduate study. Whether you plan to change direction, enhance your existing career or develop in-depth knowledge of your area of study, the University of Edinburgh provides a world-class learning experience. *Times Higher Education World University Rankings ** Research Assessment Exercise 2008

Contents Welcome to the School of Biological Sciences

Facilities and resources

Community

Employability and graduate attributes

Taught masters programmes

Online distance learning

Research at the School of Biological Sciences

Research opportunities

Funding

How to apply

Get in touch

Campus map

facebook.com/universityofedinburgh twitter.com/ApplyEdinburgh youtube.com/edinburghuniversity

www.ed.ac.uk

Thomas Jefferson American Founding Father and President (speaking to his son-in-law, Thomas Mann Randolph, as he began his studies in 1786)

Welcome to the University of Edinburgh/Contents

“You are now in a place where the best courses upon Earth are within your reach... such an opportunity you will never again have.”

Welcome to the School of Biological Sciences Our vibrant community of staff and students are continuing a 400-year history of scientific exploration with innovative work that seeks to shape tomorrow’s world. We are constantly seeking to shed light on the secrets of life through our wide-ranging research – the majority of which was ranked ‘internationally excellent’ or ‘world leading’ in the latest Research Assessment Exercise. Our commitment to pioneering science, supported by a healthy flow of grant funding, helps create an environment where world firsts are possible, for example the first genetically engineered vaccine against hepatitis B was developed here at the School of Biological Sciences. By joining us you will experience a unique opportunity to examine life processes at the very highest level.

Thriving community Our School houses about 120 principal investigators, both academic teaching staff and independently funded senior research fellows, about 200 research assistants, most of whom are postdoctoral, 170 research technicians and more than 250 PhD students.

Some 90 administrative and technical staff support the School’s academic activities.

• The Institute of Evolutionary Biology

outstanding opportunities to study your chosen field, from programmes taught at the world-famous Royal Botanic Garden Edinburgh to those that explore biological data through technology. Our students can be found discovering how to develop new products for human and animal health, or applying the latest engineering principles to the exploration of biological networks.

• The Institute of Immunology and Infection Research

Effective outcomes

Research institutes Our world-class research takes place at one of six research institutes: • The Institute of Cell Biology

• The Institute of Molecular Plant Sciences • The Institute for Stem Cell Research • The Institute of Structural and Molecular Biology Many of our researchers also participate in one or more of our numerous cross-disciplinary research centres.

Programmes Postgraduate teaching takes every advantage of our School’s expertise to give you

From producing last year’s Nexxus Young Life Scientist of the Year to offering industry with a full suite of services, the School of Biological Sciences continues to make its mark on the future. As you make your contribution, you will not only come into contact with people and opportunities in Edinburgh that will give you a distinct advantage, but you will also learn to expect an excellence in yourself and others that will keep you in demand throughout your future career.

The University of Edinburgh Biological Sciences Postgraduate opportunities Prospectus 2013 entry

Microscopy and flow cytometry are two of our major strengths. Our microscopy facilities house state-of-the-art widefield and confocal fluorescence, and transmission and scanning electron microscopes. Our flow cytometry facilities offer cutting-edge analytical and sorting capabilities.

Extensively equipped We have protein production and biophysical characterisation facilities, including surface plasmon resonance, ITC and spectroscopic equipment. We host more than 1,000m2 of controlled environment growth space and glasshouse space, and have computing resources for evolutionary biology, including four dedicated computer clusters for phyloinformatics, comparative genomics and population and quantitative trait to locus genetics.

Excellent student facilities We are based almost wholly at the University’s King’s Buildings campus, located about a 15-minute bus ride from the city centre. Students will be able to take advantage of the new Noreen and Kenneth Murray Library – named after the pioneer of the first genetically engineered hepatitis B vaccine and his late molecular geneticist wife – which opens in summer 2012, as well as the KB Learning and Teaching Cluster and the social and sports facilities at KB House. Masters students have their own dedicated common room and all postgraduate research students are given their own desk space in shared student offices.

Whatever tools you need to conduct your research, you’ll find the latest versions at the School of Biological Sciences. Our GenePool sequencing service leads the way, playing host to the NERC National Sequencing Facility and Medical Research Council Hub Facility.

Welcome to the School of Biological Sciences/Facilities and resources

Facilities and resources

www.ed.ac.uk

Community The School forms one of the largest academic groupings of biological scientists in the UK. Whether you are following a taught postgraduate programme or embarking on a research degree, you will be welcomed into a supportive and enthusiastic community of students and staff.

The MSc experience As an MSc student, there is always something to keep you motivated and inspired – from Innovative Learning Week to the Edinburgh International Science Festival. Each of our MSc programmes has a dedicated administrator who is there to help you with any queries you may have.

Research support All postgraduate research students are members of the Graduate School, which boasts an active academic and social calendar. We provide every opportunity to take part – from peer support groups and journal clubs to BioDocSoc, our society run by and for research students and staff.

dissemination of results, along with an environment for training, development and mentoring of research students and early career researchers. BioSkills, a database developed by our own researchers, makes it easy to identify fellow researchers in Edinburgh who can share their skills and expertise with you.

Links and partnerships Our participation in the work of a wide variety of interdisciplinary research centres reflects the importance we place on the collaborative approach to research. Through these connections, you’ll come into contact with researchers from varied academic backgrounds and gain insights into new approaches and techniques.

Inspiring environment

We are linked with:

Our environment of shared knowledge and expertise has led to groundbreaking and globally recognised research achievements. Our research institutes provide a forum for development of ideas, collaboration and

• MRC Centre for Regenerative Medicine

• Wellcome Trust Centre for Cell Biology • SynthSys – Centre for Synthetic and Systems Biology • Centre for Translational & Chemical Biology • Centre for Bioinformatics

• Wellcome Trust Centre for Infection, Immunity & Evolution • Edinburgh Infectious Disease Many of our research projects also involve collaborations with the Scottish Agricultural College, the Royal Botanic Garden Edinburgh, and many other academic and commercial organisations. The School of Biological Sciences is lead partner in the EASTBIO Doctoral Training Partnership, a collaboration between the universities of Aberdeen, Dundee, Edinburgh and St Andrews. Funded by the Biotechnology and Biological Sciences Research Council, EASTBIO awards 34 fully funded PhD studentships annually and provides enhanced training to its students. For further details visit www.eastscotbiodtp.ac.uk.

The University of Edinburgh Biological Sciences Postgraduate opportunities Prospectus 2013 entry

Institute for Academic Development All of our postgraduate students have the opportunity to benefit from the University’s Institute for Academic Development (IAD), which provides information, events and courses to develop the skills you will need now and in the future. The IAD offers one of the most established university research and career skills training packages in the UK. Our IAD experts will help you gain the skills, knowledge and confidence needed to move on to the next stage in your career, be that in a professional sector or within academia. The Institute provides PhD researchers and masters by research students with dedicated training in topics such as research management; personal effectiveness; communication skills; public engagement, networking and teamworking; leadership; and career management. You can gain expertise in information technology and presentation skills; confidence in undertaking independent and creative research; the ability to critically

evaluate source materials; and the capacity to construct intellectually rigorous arguments. For taught postgraduates, the IAD provides a growing range of tailored study-related and transferable skills workshops, plus online advice and learning resources. These are all designed to help you settle into postgraduate life, succeed during your studies, and move confidently onwards to the next stage of your career. By developing these broader professional skills and qualities, our postgraduate students are always in high demand. For more information please visit www.ed.ac.uk/iad/postgraduates.

Careers Service The University’s award-winning Careers Service aims to expand the horizons of all our students, enabling you to make informed career decisions in order to progress towards high personal and professional achievement and a fulfilling future, whether in work or in further study. Our goal is to offer you a world-class service.

Our teams of subject-specific expert advisers are here to help at any time during your programme of study. We offer impartial guidance and information, and can draw on relationships with a wide range of employing and training organisations. For more information on the full range of services available, including advice on starting your own business, getting published, working internationally or even volunteering, visit our website at www.ed.ac.uk/careers.

Science communication There are plenty of opportunities to develop science communication skills. The student-run BioPod series of podcasts covers stories from across the School – you might find yourself the subject of one, or helping to produce them. Similarly, our Press Gang works with the University’s Press and PR Office to spread the word on the School’s pioneering work. Many of our students are part of the EUSci team, publishing a regular science magazine, and there are also opportunities to get involved with the annual Edinburgh International Science Festival.

The School of Biological Sciences offers a research-rich environment in which to develop as scientists or entrepreneurs. However, you will learn skills that will benefit your personal and professional development regardless of the direction your career may take. The University hosts a range of services and opportunities to help you make the most of your time here.

Community/Employability and graduate attributes

Employability and graduate attributes

www.ed.ac.uk

Taught masters programmes Biodiversity & Taxonomy of Plants

Bioinformatics

www.ed.ac.uk/pg/1

www.ed.ac.uk/pg/2

MSc 1 yr FT

PGDip 9 mths FT

Programme description

The understanding of plant diversity and resources has never been more important. As we face the unprecedented challenges of climate change and environmental degradation, effective environmental surveillance and conservation depend upon detailed knowledge of plants and their habitats.

Bioinformatics is the application of computer-based approaches to understand biological processes. Modern high-throughput technologies, such as genome sequencing and microarray profiling, generate huge amounts of data.

This programme is run jointly by the University and the world-renowned Royal Botanic Garden Edinburgh (RBGEE), home to one of the world’s best living collections of plants (15,000 species across four sites, amounting to five per cent of known world species), a herbarium of three million preserved specimens and one of the UK’s most comprehensive botanical libraries. RBGE offers collections-based biodiversity research opportunities across a wide spectrum of organisms and geographical regions. This diversity, coupled with the RBGE’s world-leading research in different continents, provides an unrivalled masters programme in plant biodiversity.

Programme structure This programme consists of two terms of lectures, practicals, workshops and investigations, followed by a four-month research project. The programme includes a two-week field course in a tropical country (recently Belize). The programme is delivered mainly at RBGE but also at the University’s King’s Buildings campus. There are no optional elements to the programme – all courses are compulsory. Courses: Conservation and Sustainability; Taxonomy and Plant Collections; Biodiversity of Angiosperms; Evolution of Cryptogams and Fungi; Evolution of Angiosperms; Plant Geography; Phylogenetics and Population Genetics; Biodiversity of Cryptogams and Fungi. Your research project will be chosen in consultation with your supervisor, and will link directly with active research programmes at RBGE or other research insitutions. The field trip, together with training and a short practical exam, qualifies you for the RBGE Certificate in Practical Field Botany.

Career opportunities The programme gives you a good preparation for roles in taxonomy, while many graduates have also continued to PhD studies. Past students have entered a wide variety of jobs at research institutions, conservation agencies and elsewhere.

Minimum entry requirements A UK 2:1 degree, or its international equivalent (www.ed.ac.uk/international/ country), in biological or environmental science. Relevant work experience is desirable.

English language requirements: See page 18 Tuition fees in 2012/13* 1 yr FT: UK/EU £5,750; international £17,200 *Fees change annually. For the most up-to-date information about fees see www.ed.ac.uk/student-funding.

Programme directors: Dr Andrew Hudson/Dr Louis Ronse de Craene T: +44 (0)131 650 8651 E: btpmsc@ed.ac.uk

The aim of this MSc is to equip you with a strong background in biology, plus the computing skills and knowledge necessary to navigate the vast wealth of modern biological data. On completing this programme you will be able to take up PhD studies or bioinformatics posts in academia or in industry. The programme covers programming skills, statistical analysis and database science as well as bioinformatics. Optional courses allow students to specialise in several aspects of bioinformatics.

Programme structure The MSc comprises two semesters of taught courses followed by a research project and dissertation. The project is a key element in deciding how your career in bioinformatics should develop further. Teaching is through lectures, tutorials, seminars, computer practicals and lab demonstrations. Compulsory courses: Bioinformatics Research Proposal; MSc Dissertation (Bioinformatics). Optional courses: Applied Databases; Building a Bioinformatics Workstation; Bioinformatics 1; Human–Computer Interaction; Information Processing in Biological Cells; Introduction to Scientific Programming for MSc in Drug Discovery & Translational Biology; Molecular Modelling and Database Mining; Perl for Bioinformatics; Quantitating Drug Binding; Statistics and Data Analysis; Text Technologies; Bioinformatics Algorithms; Bioinformatics 2; Computational Systems Biology; Data Mining and Exploration; Functional Genomic Technologies; Introduction to Website and Database Design for Drug Discovery; Molecular Phylogenetics; Next Generation Genomics; Software Architecture, Process, and Management; Drug Discovery.

Research The research project is carried out independently, but under the guidance of a supervisor during the summer, with results presented in a dissertation. A wide range of projects is available through both the School of Biological Sciences and the School of Informatics.

Career opportunities The programme will give you a good preparation for further academic research or for technical or managerial roles in various commercial sectors, from medical electronics to defence.

Minimum entry requirements A UK 2:1 degree, or its international equivalent (www.ed.ac.uk/international/ country), in biological sciences. You must have a strong background in molecular biology, biochemistry or related sciences and some experience of computer science or mathematics. Students with degrees in chemistry, physics, mathematics or engineering will be considered.

Programme Director: Dr Simon Tomlinson T: +44 (0)131 651 7052 E: bioinfmsc@ed.ac.uk

The University of Edinburgh Biological Sciences Postgraduate opportunities Prospectus 2013 entry

MSc 1 yr FT PGDip 9 mths FT

Programme description Changing demographics and growing demand for food, fuel and agricultural and environmental sustainability are among the key challenges the world faces today. In this MSc you will learn research and development skills using biological principles to enable the creation of new products and services. You will investigate the economic basis for current biotechnology structures and areas of future demand, including the global pharmaceutical industry, carbon sequestration, biofuel production, innovations in drug and vaccine design and regenerative medicine. You will learn how technology can be applied to solve pressing real-world biological problems and gain the skills and expertise needed for future developments in biotechnology.

Programme structure This programme consists of two semesters of taught courses followed by a research project or industrial placement. Compulsory courses: Research Project or Industrial Placement (as available); Economics and Innovation in the Biotechnology Industry; Intelligent Agriculture; Research and Development: The Laboratory Foundation Part 1 and 2; Vaccine Development and Clinical Testing. Optional courses: Information Processing in Biological Cells; Enzymology and Biological Production; Gene Expression and Microbial Regulation; Molecular Modelling and Database Mining; Commercial Aspects of Drug Discovery; Management of R&D and Product Innovation; Building a Bioinformatics Workstation; Regenerative Medicine; Bioinformatics; Stem Cell Systems for Translational Medicine; Stem Cells for Regenerative Medicine: a Roadmap for the Clinic; Drug Discovery; Social Dimensions of Systems and Synthetic Biology.

There will be a considerable practical element to the programme. You will work in a biotechnology laboratory and will learn how experimental technology is designed and operated. Group and class projects will centre on bioelectricity, biofuels and other areas of biotechnology.

Industrial placement You can pursue a traditional laboratory-based project on campus, resulting in a dissertation, or choose to undertake an industrial placement. Support from the Scottish Funding Council has led to a new scheme enabling students to do their dissertation in a workplace setting or consultancy role, working with employers in the Scottish biotechnology sector.

Career opportunities The programme will open up a wide variety of career opportunities, ranging from sales and marketing, to research and development, to manufacturing and quality control and assurance.

Minimum entry requirements A UK 2:1 degree, or its international equivalent (www.ed.ac.uk/international/ country), with a strong background in biotechnology, biochemistry, molecular biology or related sciences.

English language requirements: See page 18 We strongly recommend that students exceed the specified minimum levels as presentations and report writing are a significant aspect of this programme.

Tuition fees in 2012/13* 1 yr FT: UK/EU £5,750; international £17,200 *Fees change annually. For the most up-to-date information about fees see www.ed.ac.uk/student-funding.

Programme Director: Professor David Arnot T: +44 (0)131 650 8643 E: biotechmsc@ed.ac.uk

www.ed.ac.uk/pg/676

Research and laboratory work

Taught masters programmes

Biotechnology

www.ed.ac.uk

Drug Discovery & Translational Biology www.ed.ac.uk/pg/3 MSc 1 yr FT PGDip 9 mths FT

Programme description Pharmaceuticals have traditionally been developed from herbal remedies or from the screening for beneficial effects of natural or synthetic chemicals. Today, the process of drug discovery is being transformed by technology. This MSc will help you explore current methods of developing drugs and therapeutic compounds for humans and animals and disease control agents for plants. You will learn about marketing, licensing and regulations, which are all part of the development process. Our multidisciplinary approach will help you link structural biology, bioinformatics, chemistry and pharmacology. You will investigate the fundamental scientific problems and techniques of drug discovery and design, alongside the challenges of developing principles for new therapeutic strategies. You will have hands-on experience of crystallographic computer programming and computation for bioinformatics. As part of the transferable skills aspect to the programme, you will consider the moral and ethical aspects of the agrochemical and pharmaceutical industries through case studies, seminars and discussions.

Programme structure This programme consists of two semesters of taught courses followed by a research project or industrial placement. Compulsory courses: Applicable Mathematics; Molecular Modelling and Database Mining; Quantitating Drug Binding; Protein Structure Determination; Commercial Aspects of Drug Discovery; Project Proposal, Literature Review and Research Project; Ethics in Science; Preparative Methods for Structural Biology Drug Discovery. Optional courses: Chemical Pharmacology; Bioinformatics 1; Functional Genomic Technologies; Synthetic Biology: Modelling; Biophysical Chemistry for MSc Drug Discovery and Translational Biology; Introduction to Scientific Programming; Medicinal Chemistry; Introduction to Website and Database Design for Drug Discovery; Detailed Characterisation of Drug or Ligand Interactions Using Surface Plasmon Resonance (SPR); Perl for Bioinformatics; Building a Bioinformatics Workstation; Bioinformatics Algorithms; Management of R&D and Product Innovation; Information Processing in Biological Cells.

Career opportunities This MSc is designed to help you pursue a career in the pharmaceutical industry or relevant government agencies, and it will provide a good background for managerial or technical roles in research, design and development. It is also a solid basis from which to continue your studies to PhD level.

Minimum entry requirements A UK 2:1 degree, or its international equivalent (www.ed.ac.uk/international/ country), in biochemistry, molecular biology, biophysics or pharmacology. Applicants with a background in chemistry, physics or computer science will also be considered. Students with other scientific or medicine backgrounds should contact the Programme Director.

English language requirements: See page 18 We strongly recommend that students exceed the specified minimum levels as presentations and report writing are a significant aspect of this programme.

Tuition fees in 2012/13* 1 yr FT: UK/EU ÂŁ5,750; international ÂŁ17,200 *Fees change annually. For the most up-to-date information about fees see www.ed.ac.uk/student-funding.

Programme Director: Dr Paul Taylor T: +44 (0)131 651 7052 E: drugdiscovery@ed.ac.uk

The University of Edinburgh Biological Sciences Postgraduate opportunities Prospectus 2013 entry

PGDip 9 mths FT

Programme structure

Specialist options within this programme are:

This programme consists of two semesters of taught courses followed by a research project.

www.ed.ac.uk/pg/6

Animal Breeding & Genetics www.ed.ac.uk/pg/763 MSc 1 yr FT (2 yrs PT available for UK/EU students) PGDip 9 mths FT

Evolutionary Genetics www.ed.ac.uk/pg/764 MSc 1 yr FT (2 yrs PT available for UK/EU students) PGDip 9 mths FT

Human Complex Trait Genetics www.ed.ac.uk/pg/765 MSc 1 yr FT (2 yrs PT available for UK/EU students) PGDip 9 mths FT

Programme description Quantitative Genetics & Genome Analysis is a suite of programmes offering specialist routes in Animal Breeding & Genetics, Evolutionary Genetics, or Human Complex Trait Genetics. The revolution in genetic mapping technology and the advent of whole genome sequences has turned quantitative genetics into one of the fastest growing areas of biology. Based in the internationally renowned Institute of Evolutionary Biology, this MSc draws from the wealth of expertise available there, as well as the teaching, research expertise and facilities of the Scottish Agricultural College, the University’s Centre for Molecular Medicine, the Medical Research Council’s Human Genetics Unit and the Roslin Institute (birthplace of Dolly the Sheep). Each year the syllabus is fine-tuned to suit current issues in evolutionary, plant, human and animal genetics.

Compulsory courses: Population and Quantitative Genetics; Genetic Interpretation; Statistics and Data Analysis; Linkage and Association in Genome Analysis; Mini Research Project; either Bioinformatics or Molecular Phylogenetics. Optional courses (selected according to degree specialisation): Quantitative Genetic Models; Molecular Evolution; Genetics of Human Complex Traits; Animal Genetic Improvement; Evolution and Climate Change.

Minimum entry requirements A UK 2:2 degree, or its international equivalent (www.ed.ac.uk/international/ country), in biological or biomedical sciences, medicine, agriculture or animal sciences, with evidence of quantitative skills. Alternatively a background in mathematics, statistics or physics with evidence of an interest in genetics.

English language requirements: See page 18 Tuition fees in 2012/13* 1 yr FT: UK/EU £5,750; international: £17,200 2 yrs PT: UK/EU £2,875 per year *Fees change annually. For the most up-to-date information about fees see www.ed.ac.uk/student-funding.

Programme Director: Professor Andrew Leigh Brown T: +44 (0)131 650 5513 E: qgen@ed.ac.uk

MSc 1 yr FT (2 yrs PT available for UK/EU students)

You will develop the in-depth knowledge and specialised skills required to apply quantitative genetics theory to practical problems, in both the biomedical and animal-science industries, and to undertake research in evolutionary genetics, population genetics and genome analysis.

Taught masters programmes

Quantitative Genetics & Genome Analysis

www.ed.ac.uk

Systems & Synthetic Biology www.ed.ac.uk/pg/649 MSc 1 yr FT PGDip 9 mths FT

Programme description We offer an opportunity to train in one of the newest areas of biology: the application of engineering principles to the understanding and design of biological networks. This new approach promises solutions to some of today’s most pressing challenges in environmental protection, human health and energy production. The MSc will provide you with a thorough grounding in the fundamentals of system and synthetic biology, ranging from understanding genome-wide data to designing and synthesising ‘Biobricks’. You will learn quantitative methods of modelling and data analysis to inform and design new hypotheses based on experimental data. The University hosts the new SynthSys Centre for Synthetic and Systems Biology, a hub for world-leading research in this area.

Programme structure The programme consists of two semesters of taught courses followed by a research dissertation, which can be a modelling or laboratory-based project. Students are advised to take our four recommended courses, which are supplemented with their choice of optional courses. Recommended courses: Information Processing in Biological Cells; Synthetic Biology: Modelling; Computational Systems Biology; Social Dimensions of Systems and Synthetic Biology. Optional courses: Neural Computation; Probabilistic Modelling and Reasoning; Molecular Modelling and Database Mining; Introduction to Java Programming; Functional Genomic Technologies; Models and Languages for Computational Systems Biology; Machine Learning and Pattern Recognition; Data Mining and Exploration; Information Theory; Perl for Bioinformatics; Regenerative Medicine; Research and Development: the Laboratory Foundation Part 1 and 2; Statistics and Data Analysis; Stochastic Models in Biology; Bioinformatics; Bioinformatics Algorithms; Biological Physics; Computational Cognitive Neuroscience; Methods for the Analysis of Networks; Molecular Phylogenetics; Next Generation Genomics; Stem Cell Systems for Translational Medicine; Bioinformatics 1; Management of R&D and Product Innovation; Drug Discovery.

Career opportunities The programme is designed to give you a good basis for managerial or technical roles in the pharmaceutical and biotech industries. It will also prepare you for entry into a PhD programme.

Minimum entry requirements A UK 2:1 degree, or its international equivalent (www.ed.ac.uk/international/ country), with a strong background in one or more of the following areas: bioinformatics, computer sciences and modelling, molecular biology and related sciences, engineering or biotechnology. Physics, mathematics or other backgrounds will also be considered.

Programme Director: Professor Peter Swain T: +44 (0)131 651 7052 E: mscssb@ed.ac.uk

The University of Edinburgh Biological Sciences Postgraduate opportunities Prospectus 2013 entry

Next Generation Drug Discovery www.ed.ac.uk/pg/766 MSc 3 yrs PT (online distance learning) PGDip 2 yrs PT

of Drug Targets; Systems Approach to Modelling Cell Signal Transduction; Modelling Metabolic Pathways; In Silico Drug Discovery; Molecular Modelling; High Throughput Drug Discovery; Commercial Aspects of Drug Discovery. Optional courses: Introduction to Biological Kinetics; Computing Skills for Drug Discovery; Chemistry for Drug Discovery; Druggable Systems.

Programme description

Career opportunities

This online programme aims to address the global problem of the falling rate at which new drugs come to market.

You will enhance your career prospects with marketable analytical and presentation skills.

You will study the challenges in developing novel drugs; the science underlying emerging fields of drug discovery; the application of new ideas to the field; how drug discovery relates to real-world health problems; the commercial aspects of drug discovery; and potential future developments. The programme offers a research-rich environment in which you can develop as a scientist and entrepreneur.

Programme structure This is a distance learning programme taught entirely online via a virtual learning environment. You will learn through a variety of teaching methods, including online tuition, peer-to-peer discussion and individual study. You will take six taught courses in each of the two first years of the MSc – a mixture of compulsory and optional courses. In your final year you pursue a research project leading to a dissertation. You can follow a single year of part-time study to qualify for a certificate, two years to achieve a diploma, and three years for the MSc. You can expect to spend about 10 hours a week on your studies. Compulsory courses: Professional Skills in Drug Discovery; Introduction to Modelling Biological Systems; Measuring Drug Binding; Structure Determination

Minimum entry requirements A UK 2:1 degree, or its international equivalent (www.ed.ac.uk/international/ country), in biotechnology, biochemistry, chemistry, medicine, molecular biology, pharmacology or related sciences.

English language requirements: See page 18 Tuition fees in 2012/13* 3 yrs PT: UK/EU £4,350 per year; international £4,350 per year *Fees change annually. For the most up-to-date information about fees see www.ed.ac.uk/student-funding.

Programme Director: Dr Paul McLaughlin T: +44 (0)131 651 7052 E: MScNGDD@ed.ac.uk

You may also be interested in: MSc Translational Medicine (Medicine & Veterinary Medicine Prospectus)

Our award-winning online learning technology enables you to interact with our highly qualified teaching staff from the comfort of your home or workplace. In fact, you will have the same access to our staff as you would if you were on campus. Our online students not only have access to Edinburgh’s excellent resources but they get the opportunity to become part of a supportive online community.

Taught masters programmes/Online distance learning

Online distance learning

www.ed.ac.uk

Research at the School of Biological Sciences Pursuing postgraduate research in the School of Biological Sciences will place you within one of the largest and most highly rated centres of its kind in the UK and one of the top 25 worldwide for life sciences*. With the majority of our research judged as either world-leading or internationally significant in the latest Research Assessment Exercise, and with a successful record of research grant applications, we can offer an environment that boasts the most up-to-date equipment and facilities to encourage research excellence and innovation. You’ll be part of an active Graduate School, numbering about 200 students, and will benefit from the support of more than 120 principal investigators, many of whom hold independent personal fellowships funded by prestigious bodies. Through our innovative * Times Higher Education World University Rankings 2011/2012

‘skills database’, you’ll have access to a broad range of expertise within our comprehensive research areas, the output of which has been assessed as second in the UK for the volume of ‘internationally excellent’ and ‘world-leading’ research.

Research routes Our School is divided into six discipline-based research institutes that cover the full spectrum of biology, from molecular structure to evolutionary and population biology. We offer two routes to a PhD: a four-year programme that includes a substantial training element and a three-year programme for students with a strong background in their chosen area.

High rate of career success We find the overwhelming majority of our students have an academic career in mind when applying for research degrees. Our latest figures show that 84 per cent of our graduates achieve this, and are now establishing

themselves in universities and other research institutions worldwide. It’s a significant achievement, one which demonstrates the quality of both our research and our standing within the international academic community. Of course, some students plan to develop a career outside academia and they are also supported.

Liam Worrall PhD Structural Biology

“I completed a PhD in Structural Biology at the University of Edinburgh. I originally joined the Walkinshaw group as a bioinformatics PhD candidate but through his supportive and motivational mentorship took up the exciting challenge of X-ray crystallography, which I have continued to investigate through a postdoctoral fellowship at the University of British Columbia, Vancouver, Canada.”

The University of Edinburgh Biological Sciences Postgraduate opportunities Prospectus 2013 entry

Cell Biology

Evolutionary Biology

www.ed.ac.uk/pg/7

www.ed.ac.uk/pg/8

PhD 3 yrs FT (6 yrs PT available for UK/EU students)

MPhil 2 yrs FT (4 yrs PT available for UK/EU students)

MSc by Research 1 yr FT (2 yrs PT available for UK/EU students)

Research environment The Institute of Cell Biology (ICB), including the Wellcome Trust Centre for Cell Biology, hosts 32 laboratories carrying world-class research into fundamental mechanisms relating to cellular function. A significant proportion of our research involves systems biology approaches and mathematical modelling.

Research environment As a research student at the Institute of Evolutionary Biology you will join the UK’s largest and most dynamic community of researchers in the field. With about 30 research groups, which includes about 50 PhD students, you’ll benefit from a busy programme of seminars, journal clubs and other research-furthering activities and will be encouraged to publish your findings. You’ll also take part in student-led courses and talks.

Our research groups comprise one of the largest communities of cell biology researchers in the UK. Many of our staff are Fellows of the Royal Society and acknowledged leaders in their field. Most PhD students who train at the ICB continue with their academic career or join research in the industry, often becoming independent group leaders.

We have an enviable record in academic career success. Our institute reflects the School-wide figure of 84 per cent for students achieving academic posts after they complete their PhD.

Diversity of interests We offer a wide scope of projects covering different areas of cell biology, including the synthesis, processing, localisation and degradation of RNA; epigenetic control of gene expression, chromosome function and genome stability; mechanisms of cell growth and duplication; the rules that govern cellular architecture; biotechnology, synthetic biology and microbial regulation. A variety of courses in microscopy, bioinformatics and proteomics are available to all our PhD students.

We go beyond the study of animal behaviour to focus on the underpinnings of the science of evolution, down to the level of DNA, to look at just how organisms fundamentally develop as a result of natural selection.

State-of-the-art facilities The ICB provides an exceptional working environment. The Wellcome Trust Centre for Cell Biology is home to the Central Optical Instrumentation Laboratory (COIL), while image analysis at the boundary of biology and physics are provided through the nearby Collaborative Optical Spectroscopy Micromanipulation & Imaging Centre (COSMIC). The ICB is proud of its cutting-edge proteomics facility and automated drug-screening platforms that facilitate discovery of new molecules that impact on cellular functions.

First-class facilities Topping our list of impressive research tools is the GenePool, one of the UK’s biggest and best university-based genomics facilities. Attracting visiting researchers from all over the UK and internationally, this next-generation facility opens up new ways of creating research data. If your study involves bioinformatics, the Ashworth Bioinformatics Support Service provides a vital link with the expertise and facilities on offer through our world-leading School of Informatics. Tuition fees in 2012/13* PhD 3 yrs FT: UK/EU £3,828; international £15,000 per year PhD 6 yrs PT: UK/EU £1,914 per year MPhil 2 yrs FT: UK/EU £3,828; international £15,000 per year MPhil 4 yrs PT: UK/EU £1,914 per year MSc by Research 1 yr FT: UK/EU £5,750; international £17,200 MSc by Research 2 yrs PT: UK/EU £2,875 per year *Fees change annually. For the most up-to-date information about fees see www.ed.ac.uk/student-funding.

www.ed.ac.uk

Tuition fees in 2012/13* PhD 3 yrs FT: UK/EU £3,828; international £15,000 per year PhD 6 yrs PT: UK/EU £1,914 per year MPhil 2 yrs FT: UK/EU £3,828; international £15,000 per year MPhil 4 yrs PT: UK/EU £1,914 per year MSc by Research 1 yr FT: UK/EU £5,750; international £17,200 MSc by Research 2 yrs PT: UK/EU £2,875 per year *Fees change annually. For the most up-to-date information about fees see www.ed.ac.uk/student-funding.

Our current research falls under three broad themes: evolutionary and quantitative genetics, evolutionary ecology and behaviour, and biodiversity and ecology. Researchers are working on everything from viral evolution and host-parasite co-evolution to natural selection in the wild and plant conservation genetics.

The School offers a wide range of research areas for postgraduate-level research, but you may also be interested in opportunities in the University’s College of Medicine & Veterinary Medicine (www.ed.ac.uk/ medicine-vet-medicine) or School of Chemistry (www.chem.ed.ac.uk).

Research at the School of Biological Sciences/Research opportunities

Research opportunities

Immunology & Infection Research

Stem Cell Research

www.ed.ac.uk/pg/488

www.ed.ac.uk/pg/11

PhD 3 yrs FT (6 yrs PT available for UK/EU students)

MPhil 2 yrs FT (4 yrs PT available for UK/EU students)

MSc by Research 1 yr FT (2 yrs PT available for UK/EU students)

Research environment Immunology and Infection Research has a proud tradition here at the School of Biological Sciences. As a researcher, you’ll be following in the footsteps of Nobel Prize winners – from malaria pioneer Sir Ronald Ross to the 2001 laureate Sir Paul Nurse – and other great names who have made groundbreaking discoveries over the years. Our leadership in infectious disease research has emerged alongside the School’s strength in population biology and quantitative genetics; our close association with biologists in these areas can be a valuable asset to your work. Our institute has more than 20 research group leaders, including nine professors. While genetics is a core discipline, our work extends over molecular and cell biology, immunology, evolutionary biology and epidemiology. Themes We cover a number of overlapping themes of study. In fundamental immunology, we look at how B and T lymphocytes interact and develop in response to antigen challenge, and how responses are initiated by dendritic cells and dampened by regulatory cells. Immune regulation of disease projects aim to design immunological interventions to ameliorate pathology or to enhance host immunity, in conjunction with new vaccines against parasites and therapies for autoimmunity. Work on molecular biology and genetics of parasites focuses on the identification of vaccine candidates and virulence factors in malaria parasites, the cell and molecular biology of trypanosomes and Leishmania parasites, and proteomics and transcriptomics of helminth worm parasites. In molecular microbiology, we look at the cell biology and post-genomic functional analysis of intracellular bacterial pathogens including Listeria and Helicobacter; and host-pathogen population biology integrates conventional immunology, pathogen research and systems-level quantitative biology, using mathematical approaches in both experimental models and the epidemiology of human and animal infections. Tuition fees in 2012/13* PhD 3 yrs FT: UK/EU £3,828; international £15,000 per year PhD 6 yrs PT: UK/EU £1,914 per year MPhil 2 yrs FT: UK/EU £3,828; international £15,000 per year MPhil 4 yrs PT: UK/EU £1,914 per year MSc by Research 1 yr FT: UK/EU £5,750; international £17,200 MSc by Research 2 yrs PT: UK/EU £2,875 per year *Fees change annually. For the most up-to-date information about fees see www.ed.ac.uk/student-funding.

Research environment Our role in the development of stem cell research began in the early 1990s, with researchers at the formerly known Centre for Genome Research producing the subject’s early publications. We grew into the first stem cell research institute in the UK, and this formed the basis of what is now a world-leading centre for multidisciplinary research in mammalian stem cell biology and regenerative medicine. As a PhD student, you’ll study in a unique environment, thanks to our location within the MRC-funded Scottish Centre for Regenerative Medicine (SCRM). This unique cross-disciplinary initiative is aimed at studying the properties and behaviour of stem cells to develop new therapies. Field of focus Research groups within SCRM cover a diverse range of areas, exploring the possibilities offered by embryonic, fetal and adult tissue stem cells. Each group focuses on a highly specific aspect of research. The themes we address include: the properties of stem cells and their niches; control of stem cell self-renewal and differentiation in vitro and in vivo; the role of stem cells in the ageing process; and the mechanisms underlying tissue degeneration and repair. The Centre’s overarching goal is to harness this knowledge to develop new therapies. We attract highly committed and motivated students and reward them with an interdisciplinary research environment that fosters achievement at the highest levels. Cutting-edge facilities You’ll be studying in the spectacular SCRM building, built in late 2011. You’ll be part of a cutting-edge PhD programme that integrates basic and clinical research, and you’ll enjoy the benefits of a shared suite of facilities and operational expertise that are second to none. A centralised cell culture facility, a transgenic animal facility, a flow cytometry service, an imaging facility and SPF animal facilities are just some of the specialised resources you’ll have at your disposal. Your colleagues will include members of the University’s College of Medicine & Veterinary Medicine who can provide a clinical perspective and as you’ll be located a stone’s throw from the Royal Infirmary of Edinburgh, you’ll also have access to hospital facilities and practising clinicians. Tuition fees in 2012/13* PhD 3 yrs FT: UK/EU £3,828; international £15,000 per year PhD 6 yrs PT: UK/EU £1,914 per year MPhil 2 yrs FT: UK/EU £3,828; international £15,000 per year MPhil 4 yrs PT: UK/EU £1,914 per year MSc by Research 1 yr FT: UK/EU £5,750; international £17,200 MSc by Research 2 yrs PT: UK/EU £2,875 per year *Fees change annually. For the most up-to-date information about fees see www.ed.ac.uk/student-funding.

The University of Edinburgh Biological Sciences Postgraduate opportunities Prospectus 2013 entry

www.ed.ac.uk/pg/10

www.ed.ac.uk/pg/12

PhD 3 yrs FT (6 yrs PT available for UK/EU students)

MPhil 2 yrs FT (4 yrs PT available for UK/EU students)

MSc by Research 1 yr FT (2 yrs PT available for UK/EU students)

Research environment With world-class facilities and lively collaborations, our Institute of Molecular Plant Sciences provides you with the ideal environment in which to pursue postgraduate research. We’re the most active university department for plant sciences in the UK, hosting one of the country’s few plant-based research and teaching institutes. Our 14 research groups each make vital contributions to new discoveries. You’ll be able to choose from a broad range of topics, applying multidisciplinary approaches – from computational biology to molecular genetics and cell biology and to subjects ranging from plant growth, development and evolution to plant pathogen interactions. Broad scope We offer a wide range of research interests, including developmental biology; speciation and natural variation (making use of our strong connections with the Royal Botanic Garden Edinburgh); circadian clocks and photobiology; bio-imaging; biochemistry and cell biology; and molecular plant pathology. Investment in facilities Our institute was recently refurbished and since then, substantial investment has been made in ensuring the equipment and facilities you’ll use are as good as those anywhere in the world. You’ll work in purpose-built laboratories and specialist facilities for plant growth, including a high-throughput Arabidopsis genetics facility, a suite of biochemistry equipment, more than 700m2 of controlled environment growth rooms and 2,200m2 of glasshouses, including a new Category 3 containment glasshouse. Our strong links with SynthSys, the Centre for Synthetic and Systems Biology and the Royal Botanic Garden Edinburgh give you access to an even broader range of facilities and expertise, as do our associations with the Scottish Crop Research Institute. Tuition fees in 2012/13* PhD 3 yrs FT: UK/EU £3,828; international £15,000 per year PhD 6 yrs PT: UK/EU £1,914 per year MPhil 2 yrs FT: UK/EU £3,828; international £15,000 per year MPhil 4 yrs PT: UK/EU £1,914 per year MSc by Research 1 yr FT: UK/EU £5,750; international £17,200 MSc by Research 2 yrs PT: UK/EU £2,875 per year *Fees change annually. For the most up-to-date information about fees see www.ed.ac.uk/student-funding.

Research environment Closely allied with the Institute of Cell Biology, our group studies the structure and impact of biomolecules, from their atomic structure and assembly into molecular machines, to studying the large-scale effects of these molecular assemblies. We offer a combination of world-class facilities and training that sets us apart as one of the premier centres for this type of research in the UK. You’ll be supported by well-published staff who are leaders in their field and will benefit from a comprehensive programme of seminars, symposiums, distinguished visitors and social events. Finding answers Our groups investigate molecular structure using techniques such as biomolecular nuclear magnetic resonance, X-ray crystallography, computational structure prediction and cryo-electron microscopy. We also cover the full panoply of methods for protein production and biological characterisation. A growing area of our research is the development of technologies that combine chemical library generation with proteomics and high-speed imaging methods to allow identification of new ligand-protein interactions. Through links with medicine and veterinary medicine, we aim to make a real impact on healthcare through drug discovery projects. Outstanding facilities Your work will benefit from our strong links with a number of on-site research centres, including the Wellcome Trust Centre for Cell Biology, the Centre for Science at Extreme Conditions, the Centre for Translational and Chemical Biology, and SynthSys, the Centre for Synthetic and Systems Biology. We offer outstanding laboratories, equipment and training, particularly in the areas of protein production and characterisation, cryo-EM and macromolecular X-ray crystallography. Tuition fees in 2012/13* PhD 3 yrs FT: UK/EU £3,828; international £15,000 per year PhD 6 yrs PT: UK/EU £1,914 per year MPhil 2 yrs FT: UK/EU £3,828; international £15,000 per year MPhil 4 yrs PT: UK/EU £1,914 per year MSc by Research 1 yr FT: UK/EU £5,750; international £17,200 MSc by Research 2 yrs PT: UK/EU £2,875 per year *Fees change annually. For the most up-to-date information about fees see www.ed.ac.uk/student-funding.

Structural & Molecular Biology

Research opportunities

Molecular Plant Sciences

www.ed.ac.uk

Funding A large number of scholarships, loans and other funding schemes are available for your postgraduate studies. You can find the full range at www.ed.ac.uk/student-funding. Awards are offered by the School of Biological Sciences, the College of Science & Engineering, the University of Edinburgh, the Scottish, British and international governments and funding bodies. Below we list a selection of potential sources of financial support for postgraduate students applying to the School of Biological Sciences. Where an entry does not include a web address, please use the web link above to navigate to the particular award.

University of Edinburgh scholarships

Other sources of funding

China Scholarships Council/University of Edinburgh Scholarships

Commonwealth Scholarships

A number of scholarships for PhD study to candidates who are citizens and residents of China. www.ed.ac.uk/student-funding/china-council

College of Science & Engineering The College of Science & Engineering has funds from endowments and donations that are available to provide funding for studentships.

Edinburgh Global Masters Scholarships A number of scholarships available to international students for masters study. www.ed.ac.uk/student-funding/masters

Edinburgh Global Research Scholarships These scholarships are designed to attract high-quality international research students to the University. www.ed.ac.uk/student-funding/global-research

Edinburgh Santander Masters Scholarships Several scholarships are available to students from a number of countries for masters study. www.ed.ac.uk/student-funding/santander

Edinburgh UK/EU Masters Scholarships Scholarships for UK and EU students who have been accepted on a full-time masters degree programme. www.ed.ac.uk/student-funding/uk-masters

For students who are resident in any Commonwealth country, other than the UK. www.dfid.gov.uk/cscuk

Fulbright Scholarships Scholarships open to US graduate students in any subject wishing to study in the UK. www.iie.org/fulbright

Marshall Scholarships Open to outstanding US students wishing to study at any UK university for at least two years. www.marshallscholarship.org

Scotland’s Saltire Scholarships A number of scholarships open to citizens of Canada, China, India and the US, undertaking masters-level study in Scotland. www.ed.ac.uk/student-funding/saltire

The University of Edinburgh Graduate Discount Scheme We offer a 10 per cent discount on postgraduate fees for all alumni who have graduated with an undergraduate degree from the University. www.ed.ac.uk/student-funding/discounts

Principal’s Career Development PhD Scholarships A number of awards, open to UK, EU and international PhD students. www.ed.ac.uk/student-funding/development

Principal’s Indian Masters Scholarships 15 scholarships are available to students from India for masters study. www.ed.ac.uk/student-funding/masters-india

School of Biological Sciences Research Scholarships Within the School of Biological Sciences, a number of highly competitive research scholarships are available each year to new postgraduate research students. Further details are available on the School website. www.ed.ac.uk/schools-departments/biology/postgraduate/fees-funding

Robert Starr MSc High Performance Computing Scotland’s Saltire Scholarship

Wellcome Trust Four-Year PhD Programme in Cell Biology Awarded to five outstanding graduates per year, this programme provides experience of advanced techniques in cell biology in an excellent research environment. www.wcb.ed.ac.uk/phd

“The Scottish Government’s initiative to attract international students from Canada, China, India and the US through the Saltire Scholarship Scheme as well as the University of Edinburgh’s help and support for international students has helped provide me with an opportunity that I would never have conceived prior to starting my studies at Edinburgh.”

The University of Edinburghâ&#x20AC;&#x201A; Biological Sciences Postgraduate opportunities Prospectus 2013 entry

Normally only those UK/EU students who have been resident in the UK for the preceding three years are eligible for a full award. For some awards, candidates who are EU nationals and are resident in the UK may be eligible for a fees-only award. www.ed.ac.uk/student-funding/research-councils

The Canada Student Loans Program The University is eligible to certify Canadian student loan applications. Full details on eligibility and how to apply can be found online. www.ed.ac.uk/student-funding/canadian-loans

The Student Awards Agency for Scotland This department of the Scottish Government has offered loans to postgraduate students in the past. At time of going to press arrangements for study in 2013/14 were under review. www.saas.gov.uk

US Student Loans The University is eligible to certify loan applications for US loan students. Full details on eligibility and how to apply can be found online. www.ed.ac.uk/student-funding/us-loans

Research councils offer awards to masters, MPhil and PhD students in most of the Schools within the University of Edinburgh. All studentship applications from the research councils must be made through the University, through your School or College office. Awards can be made for both taught and research programmes.

Financial aid

Funding

Research council awards

www.ed.ac.uk

How to apply Taught programmes For taught masters programmes, you will in most cases need a UK degree with a minimum 2:1 classification, or its international equivalent, generally in a related area. It is expected that you will have a good understanding of the field you propose to study, and at least some relevant work experience. Entry requirements for some programmes may vary, so check the details for the specific programme you wish to apply for. To apply for one of our taught masters programmes, simply visit www.ed.ac.uk/pg/ degrees and make your application online. It’s a straightforward process, and you’ll be able to set up an online account, which lets you save your application and continue at another time.

its international equivalent) and clear evidence of interest in and suitability for research with one of our supervisors. We interview all shortlisted applicants.

Joining us from overseas

If you are interested in applying for a research degree, your first step is to identify your area of interest and explore the research possibilities within that group. Visit www.ed.ac.uk/ biology/phdproj.

about their visa options and our Integrated

You should also discuss your proposal with potential supervisors. Visit www.ed.ac.uk/ biology/people. UK students should see also the PhD projects available through the EASTBIO Doctoral Training Partnership. Visit www.eastscotbiodtp.ac.uk. The final step is to formally apply through our online admissions system. Go to www.ed.

If you’re considering a particular programme and have questions, please contact the postgraduate team within our Biology Teaching Organisation to discuss them with you. See Get in touch, page 20.

ac.uk/pg/degrees and follow the prompts.

Research programmes

research degrees, visit our Graduate School

We encourage applications from UK, EU and international applicants to our research programmes. Competition for funded places is fierce so please ensure you apply early; our first application deadline is in December each year. Your application will be judged on academic excellence (in most cases we expect a 2:1 or

postgraduate.

The system will allow you to set up an account so that you can save your application and revisit it at another time.

International applicants are advised to check the University’s website to find out more English for Academic Purposes (IEAP) programme. Please visit www.ed.ac.uk/ international/ieap.

International agents The University has certified representative agents in the following locations: Brunei, Canada, China, Gulf Region, Hong Kong, India, Japan, Jordan, Korea, Malaysia, Mexico, Nigeria, Norway, Russia, Saudi Arabia, Singapore, South Africa, South Korea, Taiwan, Thailand, Turkey, Zambia and Zimbabwe. International applicants can use an agent to help guide them through the application process if necessary. Contact details for all our agents can be found at www.ed.ac.uk/studying/ international/agents.

For more information on postgraduate website at www.ed.ac.uk/biology/

Alternatively, we are happy to talk to you about any specific enquiries. See Get in touch, page 20.

English language requirements Students whose first language is not English must show evidence of one of the qualifications in the box below.

IELTS 6.5 (with no score lower than 6.0 in each section) TOEFL iBT Total 92 (with no score lower than 20 in each section). Pearson Test of English 61 (with no score lower than 56 in each section) Cambridge CPE Grade B

Please note: • English language requirements can be affected by government policy so please ensure you visit our website for the latest details. www.ed.ac.uk/english-requirements/pg/cse • Your English language certificate must be no more than two years old at the beginning of your degree programme. • A degree from an English-speaking university may be accepted in some circumstances. • Cambridge tests are accepted only for applicants who do not need Tier 4 visas to enter the UK.

Cambridge CAE Grade A Abbreviations: IELTS − International English Language Testing System; TOEFL iBT − Test of English as a Foreign Language Internet-Based Test; CPE − Certificate of Proficiency in English; CAE − Certificate in Advanced English

The University of Edinburgh Biological Sciences Postgraduate opportunities Prospectus 2013 entry

www.ed.ac.uk

“I gained a lot from my postgrad research experience at the School of Biological Sciences. The School contains many research groups that often collaborate on big projects. Students with different backgrounds and skills (from wet lab scientists to software programmers) combine their efforts to advance further in a field that is becoming more interdisciplinary every day. My supervisors were always available for help and their advice was aimed at opening the door for more possibilities, keeping me always on edge and thinking ahead. The School was clearly focused on helping students and I truly believe that the skills of critical thinking and teamwork, along with a renewed independence, determination, and multicultural awareness, couldn’t have been better gained anywhere else than at Edinburgh.”

PhD Computational Biology

How to apply

Wissam Mehio

Get in touch Contact us

For more information about our postgraduate research programmes, please contact:

For more information about taught MSc programmes, please contact:

Dr Caroline Proctor

Vicky Mactaggart Biology Teaching Organisation G5, Darwin Building Kingâ&#x20AC;&#x2122;s Buildings Mayfield Road Edinburgh, UK EH9 3JR T: +44 (0)131 651 7052 E: pgtbiol@ed.ac.uk For enquiries relating to specific programmes, please refer to the contact details provided for each programme listing.

Graduate School of Biological Sciences 117, Darwin Building Kingâ&#x20AC;&#x2122;s Buildings Mayfield Road Edinburgh, UK EH9 3JR T: +44 (0)131 650 5327 E: gradbiol@ed.ac.uk To discuss your PhD proposal, you should identify potential supervisors at www.ed.ac.uk/biology/people.

Visit us Our postgraduate Open Day is your opportunity to come and meet current staff and students. Our next campus-based Open Day takes place on Friday 23 November 2012. For further details, please visit www.ed.ac.uk/postgraduate-open-day. We also run online information sessions for prospective postgraduate students throughout the year. To find out more, visit www.ed.ac.uk/pg/open-day/online-events.

The University of Edinburgh Biological Sciences Postgraduate opportunities Prospectus 2013 entry

LONDON RD

Y RR

S. BR IDGE

TT PO

E ST

HOM

ARGYLE PL

MEL VILLE DR

MELVILLE TER

A1 SOUTH

CHURC H LANE

EST DW OA NR O ST ING DD DU

POLLOCK HALLS

SCIENNES RD

COMMONWEALTH POOL

ROYAL HOSPITAL FOR SICK CHILDREN

D IEL YF MA

RIE MAINS RD NIDD

RD DY LA

RD TLE CAS

PK AR

WE ST M AINS RD

LL MI AIG CR

R TE

D RD FIEL MAY

NUE AVE ORD CKF BLA

THE ROYAL OBSERVATORY

E VIL SA ST WE

R ILLA IGM CRA

NS GD

D ON R RAST KILG

STUDENT RECRUITMENT AND ADMISSIONS AND INTERNATIONAL OFFICE

D HR EIT LK DA

AD GE RO GR AN

ST TO MIN

MARCHMONT RD

CLUNY GDNS

ST ESTON E.PRESTON ST W.PR

IDE AYS SEW CAU

N LOA USE HO ITE WH

N ROAD THEAR STRA

MAIN LIBRARY

D R PK R NDE RRE WA

SCOTTISH PARLIAMENT

H RD KEIT AL

THE MEADOWS

HOLYROOD PARK

N SO OL NIC T HS

GE GEOR RE SQUA

MCEWAN HALL

THE UNIVERSITY OF EDINBURGH VISITOR CENTRE

C LEU CC BU

EDINBURGH DENTAL INSTITUTE

HOLYROOD PALACE

A NG NO

CENTRE FOR SPORT & EXERCISE

OLD COLLEGE

W RO ER

L L AURISTON P

EDINBURGH COLLEGE OF ART

PLEASANCE

IDGE

T KE AR GRA SSM

R LY HO

M OYAL ST (R

ATE COWG S ST BER M CHA

E IV BR

MORRISON ST

AW SO

GEORG

N LOTHIA

LA DY L

IDGE

UND MO THE

N. BR

ST T

N ER

HIGH

EDINBURGH CASTLE

C OA PR

NEW COLLEGE

ILE)

W ILL OW BR AE RD

OL D

W ND

A SH

WAVERLEY TRAIN STATION

CES PRIN

RD BELLO PORTO

REG ENT TER MORAY HOUSE SCHOOL OF EDUCATION

S RGE

GEO

CITY OBSERVATORY

ITH

T EN S

QUE

CALTON HILL

ST ANDREW’S BUS STATION

TO THE WESTERN GENERAL HOSPITAL

UNIVERSITY OF EDINBURGH PLAYING FIELDS PEFFERMILL OL DD ALK EIT HR D A7

D TR ON LEM ESS

ROYAL INFIRMARY OF EDINBURGH & UNIVERSITY OF EDINBURGH MEDICAL SCHOOL (LITTLE FRANCE)

UNIVERSITY KING’S BUILDINGS

ON ST NG KI

E AV

A7 SOUTH

N TO ER LIB

AE BR 01 A7

ALNWICKHIL L RD

LIBERTO N DRIVE

A702 SOUTH A701 SOUTH THE ROYAL (DICK) SCHOOL OF VETERINARY STUDIES (EASTER BUSH)

University building

Photography by Paul Dodds Yao Hui Andrew Lee Norrie Russell Laurence Winram Printed by J Thomson Colour Printers www.jtcp.co.uk

www.ed.ac.uk

Published by Communications and Marketing The University of Edinburgh

Graduate School of Biological Sciences

A90 ROUTE TO FORTH ROAD BRIDGE AND NORTH

Get in touch/Campus map

Campus map

Postgraduate Open Day: 23 November 2012

Semester 1: 16 September–20 December 2013

Induction Week: 9–13 September 2013

Semester 2: 13 January–23 May 2014

FSC This publication is available online at www.ed.ac.uk and can be made available in alternative formats on request. Please contact Communications.Office@ed.ac.uk or call +44 (0)131 650 2252.

EDINB E56

© The University of Edinburgh 2012. No part of this publication may be reproduced without written permission of the University. The University is a charitable body, registered in Scotland, with registration number SC005336.