The Diptera of Suffolk. Part 2: Bibionoidea and Sciaroidea by Suffolk Naturalists' Society

SUFFOLK DIPTERA CHECKLIST

THE DIPTERA OF SUFFOLK PETER VINCENT

Part Two: Bibionidae, Bolitophilidae, Diadocidiidae, Ditomyiidae, Keroplatidae, Mycetophilidae, Sciaridae, Cecidomyiidae. The checklist begins with the first infraorder of the Lower Diptera the Tipulomorpha. In Part Two this is followed by the second infraorder of the Lower Diptera the Bibionomorpha. The Bibionomorpha contains two superfamilies, the Bibionoidea and the Sciaroidea. The Bibionoidea contains only one family, Bibionidae, whereas, the Sciaroidea contains seven families, Bolitophilidae, Diadocidiidae, Ditomyiidae, Keroplatidae, Mycetophilidae, Sciaridae, and Cecidomyiidae. The Bibionidae are robust, hairy flies and some 215mm in length. The body colour is usually black but is partially red or yellow in some cases. The family includes the largest Bibionid Bibio marci (Linnaeus, 1758), the familiar St Markâ&#x20AC;&#x2122;s fly. In Suffolk the Bibionidae are reasonably well known and of the 18 British species 17 have been recorded in the county. Bibio marci Trans. Suffolk Nat. Soc. 55 (2019)

B. Heather

Introduction The records that form this checklist are from the Watsonian vice-counties of East (v.c. 25) and West (v.c. 26) Suffolk which are of greater extent than the modern administrative county boundaries. Diptera are not well recorded in Suffolk, with few records of even quite common species. Therefore, although records are in the main collected at monad (O.S.1km square) or greater accuracy, in order to assess the relative abundance and distribution of a diptera species the hectad (O.S.10km square) is used as the base recording unit. Suffolk comprises of 54 hectads or part hectads. In this checklist of Suffolk Diptera, the arrangement of families follows as set out in Chandler (1998) and updates (Dipterists Forum, 2019). The classification of British diptera starts with the Lower Diptera (Nematocera) followed by the Brachycera. Within families, all taxa are listed alphabetically. Altogether there are at present 107 families of diptera recognised within the British fauna, and the British Isles list of diptera currently (June 2019) runs to some 7187 species (of which 41 are recorded from Ireland only) (Chandler, 2019), The layout of the checklist is that for each family the first division is by genus and then by species in alphabetical order. For most families, subfamilies and tribes are also recognised: and the genus can also be divided at subgenus level. For every individual species the species name (in bold type) is followed by the authority, then details of the first record of that species from Suffolk - place, date (to as much accuracy as possible) and recorder (where known). To give an indication of the distribution and the abundance of each species, this is followed by a list of all the hectads each species has occurred and the number of known records (in square brackets).

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The families of Bolitophilidae, Diadocidiidae, Ditomyiidae, and Keroplatidae were all, until recently, subfamilies contained within the Mycetophilidae. However, revisions by European authors now consider them as a series of families in their own right. This group of families together with the Mycetophilidae, are the ‘true’ fungus gnats. In appearance they appear hump-backed, with long coxae and well-developed tibial spurs. They are generally some combination of brown, black and yellow, rarely conspicuously patterned, but sometimes brightly coloured like Hymenoptera. Most species are rather small, but some can reach a length of 10mm. There are 16 British Bolitophilidae species, at present 7 of which have been found in Suffolk, 3 British Diadocidiidae - 2 from Suffolk, 3 British Ditomyiidae - 2 from Suffolk, 52 British Keroplatidae - 28 from Suffolk and 493 British Mycetophilidae - 268 from Suffolk. The Sciaridae have similar appearance to the fungus gnats described above, however, they are readily separable in having the eyes produced over the antennae forming an eye-bridge and in having shorter coxae. They are generally dark in appearance and therefore are sometimes referred to as ‘black fungus gnats or midges’. Sciarids are often very small, rarely exceeding 4mm in length, lacking strong bristles; their wings have a characteristic venation but are unmarked. There are 266 species on the British checklist, but only 69 of these have been recorded from Suffolk. Bolitophilidae, Diadocidiidae, Ditomyiidae, Keroplatidae, Mycetophilidae, and Sciaridae are not well recorded in Suffolk. Identification issues mean that these families are often only recorded by specialists. This is especially true for the Sciaridae where all known reliable records were published in one paper (Menzel, Smith & Chandler, 2006). The majority of other fungus gnat records used in this checklist have been kindly supplied by Peter Chandler - organiser of the Dipterists Forum Fungus Gnat Recording Scheme. The Cecidomyiidae are very fragile, small long-legged insects, usually only 2–3 mm in length and many are smaller. They are characterised by long slender antennae with bead-like segments bearing a circlet of small spines and by hairy wings, unusual in Diptera. They are commonly known as gall midges as most, but not all of them, induce gall formation. Galls are caused by the larvae of gall midges feeding within plant tissue, creating abnormal plant growths. Adult Cecidomyiids are difficult to identify both at the larval and adult stages. However, the galls that are produced are unique to each species and are readily identifiable. The study of gall midges was originally undertaken by botanists rather than entomologists and even today most recording is carried out by specialist cecidologists. The arrangement of the Cecidomyiidae subfamilies in the checklist is the usual, well established, order based on phylogenetic relationships opposed to the standard alphabetical listings. As it stands the Cecidomyiidae are the largest family within the British diptera with some 654 species and, of those 97 have been recorded in Suffolk. All the Suffolk Cecidomyiidae are associated with higher plants, or with other mostly plant associated organisms. The host, where known, of the first known county record is noted in the checklist. Many of the gall midge records used in this checklist have been kindly provided by Jeremy Bowdrey - Suffolk plant gall recorder.

Trans. Suffolk Nat. Soc. 55 (2019)

SUFFOLK DIPTERA CHECKLIST

Abbreviations used in the checklist: TSNS - Transactions of the Suffolk Naturalists’ Society, BPGS - British Plant Gall Society. References Chandler, P.J. (ed.) (1998). Checklists of Insects of the British Isles (New Series). Part 1: Diptera. Handbooks for the Identification of British Insects 12(1): 1–234. Chandler, P.J. (2019). Corrections and Changes to the Diptera Checklist (41). Dipterists Digest (Second Series) 26: 33–34. Dipterists Forum. (2019). An Update of the 1998 Checklist of Diptera of the British Isles Dipterists Forum [online] http://www. dipterists.org.uk/checklist/ [Accessed 11 November 2019] Menzel, F., Smith, J.E. & Chandler, P.J. (2006). The sciarid fauna of the British Isles (Diptera: Sciaridae), including descriptions of six new species. Zoological Journal of the Linnaean Society 146: 1–147. Suffolk Diptera Checklist (Part Two)

BIBIONIDAE BIBIO Geoffroy, 1762 anglicus Verrall, 1869 - Icklingham, 28 May 1929, C. Morley TL 77,86,96 TM 03,07,14,26,46,57 [18] clavipes Meigen, 1818 - Foxhall, August 1895, C. Morley TL 67,78 TM 07,13,14,24,35 [6] ferruginatus (Linnaeus, 1767) - Redgrave, 1978, P. Withers TM 07 [1] hortulanus (Linnaeus, 1758) - Ipswich, 20 May 1896, R. Bennett TL 76,88 TM 03,07,14,46,57 [6] johannis (Linnaeus, 1767) - Foxhall, 20 April 1895, C. Morley TL 88,94 TM 04,07,14,24,27,35,37,45,46,48 [17] lanigerus Meigen, 1818 - Monks Park Wood, Bradfield St Clare, 1901, C. Morley TL 77,78,87,95 TM 07,24,46,47,48 [10] leucopterus (Meigen, 1804) - Monk Soham, April 1940, C. Morley TL 87,95 TM 05,26,46 [5] longipes Loew, 1864 - Ipswich, 24 Oct 1895, C. Morley TL 86,95 TM 13,14,46,57 [6] marci (Linnaeus, 1758) - Monk Soham, 7 May 1909, C. Morley TG 40,50 TL 77,78,84,87,93,94,95,96,97 TM 03,05,07,13,14,15.17.24.25,27,36,37,39,44,46,47,57,59 [57] nigriventris Haliday, 1833 - Tuddenham Fen, 1916, C. Morley TL 77,87 TM 05,07,24,46 [7] pomonae (Fabricius, 1775) - Southwold, 20 July 1901, C. Morley [1] no recent records reticulatus Loew, 1846 - Raydon Woods, 1941, C. Morley TL 87,95 TM 03 [3] varipes Meigen, 1830 - Ipswich, 1899, E.W. Platten TL 87,88,94 TM 07,14,46 [6]

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DILOPHUS Meigen, 1803 bispinosus Lundström, 1913 - Wangford Wood, 13 September 1912, C. Morley TM 35,47 [2] no recent records febrilis (Linnaeus, 1758) - Pakefield Cliffs, August 1908, C. Morley TL 78, 95 TM 03,14,23,34,37,38,44,46 [21] femoratus Meigen, 1804 - Wortham Ling, 9 June 1900, C. Morley TL 88 TM 07,13,14,23,27,28,45,46,47 [20] humeralis Zetterstedt, 1850 - Felixstowe, 19 May 1897, C. Morley TL 76 TM 24,33 [3]

BOTOPHILIDAE BOLITOPHILA Meigen, 1818 Subgenus BOLITOPHILA sensu stricto cinerea Meigen, 1818 - Cavenham Heath, 30 April 1994, I. Perry TL 77,78,87,95 TM 34,35,45,46,47 [23] saundersii (Curtis, 1836) - Bradfield Woods, 16, July 1990, B.R. Laurence TL 77,78,87,95 TM 34,35,45,46,48 [33] Subgenus CLIOPISA Enderlein, 1936 glabrata Loew, 1869 - Blythburgh Wood, 14 September 1912, C. Morley TL 77,87,95 TM 47 [6] hybrida (Meigen, 1804) - Orford, 21 August 1908, J.J.F.X. King TL 77,78,87,95 TM 15,44 [7] modesta Lackschewitz, 1937 - The King’s Forest, 28 October 2001, I. Perry TL 77,87,88 95 [7] occlusa Edwards, 1913 - The King’s Forest, 29 May 2000, I. Perry TL 86,87,95 [10] pseudohybrida Landrock, 1912 - The King’s Forest, 9 October 1994, I. Perry TL 77,78,87,95 TM 24,35 [12]

DIADOCIDIIDAE DIADOCIDIA Ruthe, 1831 Subgenus DIADOCIDIA sensu stricto ferruginosa (Meigen, 1830) - Orford, 8 July 1908, J.J.F.X. King TL 77,77,86,87 TM 24,34,35,44,46,47 [26] spinosula Tollet, 1948 - Scotland Fens, Boyton, 26 October 1996, P.J. Chandler TL 78,87,95 TM 34 [9]

DITOMYIIDAE DITOMYIA Winnertz, 1846 fasciata (Meigen, 1818) -Tostock, May 1898, W.H Tuck TL 78,87,95,96 TM 46 [9] SYMMERUS Walker, 1848 annulatus (Meigen, 1830) - Barham, 22 June 1996, P.J. Chandler TL 87,88,95 TM 04,47 [6]

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KEROPLATIDAE KEROPLATINAE Keroplatini CEROTELION Rondani, 1856 striatum (Gmelin, 1790) - Monk Soham,1910, C. Morley TL 87 TM 26,59 [4] KEROPLATUS Bosc, 1792 testaceus Dalman, 1818 - Staverton Thicks, Wantisden, 24 June 1903, C. Morley TM 26,35 [2] no recent records Orfeliini ASINDULUM Latreille, 1805 nigrum Latreille, 1805 - Henstead, 12 July 1900, C. Morley TL 77 TM 16,39,48 [5] no recent records ISONEUROMYIA Brunetti, 1912 semirufa (Meigen, 1818) - Bildeston, 30 July 1898, C. Morley TL 77,78,86,87,94 TM 26,35,44,48,57 [13] MACRORRHYNCHA Winnertz, 1846 flava Winnertz, 1846 - Monk Soham, June 1920, C. Morley TL 66,87,95 TM 07,26,44,45 [7]

MONOCENTROTA Edwards, 1925 lundstroemi Edwards, 1925 -Thelnetham, 8 September 1940, C. Morley TM 07,47 [2] NEOPLATYURA Malloch, 1928 flava (Macquart, 1826) - Bradfield Woods, 16 July 1990, B.R. Laurence TL 95 [1] modesta (Winnertz, 1863) -The King’s Forest, 6 August 1994, I. Perry TL 87,88 [2] nigricauda (Strobl, 1893) - Orford, 10 July 1908, J.J.F.X. King TL 87 TM 44 [3] ORFELIA Costa, 1857 discoloria (Meigen, 1818) - Orford, July-August 1908, J.J.F.X. King TL 87 TM 44 [2] fasciata (Meigen, 1804) -The King’s Forest, 8 July 1990, I. Perry TL 87,95 TM 47 [14] earlier records in TSNS without details lugubris (Zetterstedt, 1851) -The King’s Forest, 24 June 1989, I. Perry TL 76,77,78,97 [9] nemoralis (Meigen, 1818) - Monk Soham, June 1920, C. Morley TL 66,87,95 TM 25,44,45,46,47,57 [13] nigricornis (Fabricius, 1805) -Westleton, 21 June 1998, D.A. Smith TM 34,47 [2] ochracea (Meigen, 1818) - Orford, 10 July 1908, J.J.F.X. King TL 78,958 TM 04,07,44 [12] pallida (Staeger, 1840) - Bromeswell, 10 June 1990, P.J. Chandler TL 95 TM 34,47 [3] PLATYURA Meigen, 1803 marginata Meigen, 1804 - Ipswich, 1 June 1897, C. Morley TL 77,94,95 TM 14,48 [7]

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PYRATULA Edwards, 1929 perpusilla (Edwards, 1913) - Boyton, 19 June 1908, G.H. Verrall TL 88 TM 34 [2] zonata (Zetterstedt, 1855) - Monk Soham, July 1915, C. Morley TL 77,87,95 TM 26 [5] URYTALPA Edwards, 1929 dorsalis (Staeger, 1840) - Ampton, 1912-1916, C.G. Nurse TL 86,87,88 TM 26,46 [10] MACROCERINAE MACROCERA Meigen, 1803. anglica Edwards, 1925 - Orford, July-August 1908, J.J.F.X. King TL 87,95 TM 04,26,44 [5] angulata Meigen, 1818 - Orford, July-August 1908, J.J.F.X. King TL 66,77,87,95 TM 24,44,58 [11] centralis Meigen, 1818 - Orford, July-August 1908, J.J.F.X. King TL 88,87,95 TM 07,16,26,44,45 [15] crassicornis Winnertz, 1863 - Monk Soham, 17 August 1930, C. Morley TL 86,88 TM 26 [3] fasciata Meigen, 1804 - Shrubland Park, Barham, 27 August 1920, C. Morley TL 65,77,78, 86,87,95 TM 15,26,58 [15] lutea Meigen, 1804 - Frostenden, 8 June 1977, A.M. Hutson TL 87 TM 48 [2] earlier records in TSNS without details maculata Meigen, 1818 -Timworth, 1924, C.G. Nurse TL 77,84,86 [3] nigricoxa Winnertz, 1863 - Mildenhall, 13 July 2003, P.J. Chandler TL 77,84,86,95 [12] parva Lundstrรถm, 1914 - Bromeswell, 10 June 1990, P.J. Chandler TM 34 [1] phalerata Meigen, 1818 - Orford, July-August 1908, J.J.F.X. King TL 66,77,78,84,86,87,95 TM 04,24,26,35,37,44 [24] stigma Curtis, 1837 - Staverton Thicks, Wantisden, 14 June 1904, C. Morley TL 77,95 TM 07,13,14,24,26,33,35,37,44 [16] stigmoides Edwards, 1925 - Newmarket, 23 September 1888, G.H. Verrall TL 66,67,77,78,87 TM 07,15,24,34,35,44,46,47,58 [30] vittata Meigen, 1830 -Tuddenham Fen, 21 June 1909, C. Morley TL 77,78,86,87,88,95 TM 24,26,34,46,47,48 [39]

MYCETOPHILIDAE GNORISTINAE APOLEPHTHISA Grzegorzek, 1885. subincana (Curtis, 1837) - Staverton Thicks, Wantisden, 24 April 1952, J.E. Collin TL 77,78,86,87,94 TM 34,35,47 [16] BOLETINA Staeger, 1840 basalis (Meigen, 1818) - Bentley Woods, 13 May 1900, C. Morley TL 95 TM 13 [2] dubia (Meigen, 1804) - Flatford, 18 July 1951, L. Parmenter TL 87, TM 03,24,47,59,87 [5] gripha Dziedzicki, 1885 - Benacre Broad, 11 August 1974, A.M. Hutson TL 77,87,95 TM 35,43,46,47,58 [29] Trans. Suffolk Nat. Soc. 55 (2019)

SUFFOLK DIPTERA CHECKLIST

griphoides Edwards, 1925 - Walberswick, 19 April 2018, I. Perry, TL 87 TM 47 [2] nitida Grzegorzek, 1885 - Cavenham Heath, 14 May 1994, I. Perry TL 77 [1] pallidula Edwards, 1925 - Benacre Broad, 11 August 1974, A.M. Hutson TM 46,58 [2] plana Walker, 1856 - Bromeswell, 10 June 1990, P.J. Chandler TL 87,88,95 TM 24,34 [8] sciarina Staeger, 1840 - Dunwich Forest, 18 October 1996, P.J. Chandler TL 78,87,88 TM 34,44,45,47,59 [12] trispinosa Edwards, 1913 - The King’s Forest, 20 May 2002, I. Perry TL 87 [5] trivittata (Meigen, 1818) - Redgrave Fen, 25 May 1974, A.M. Hutson TL 77,78 TM 07,24,34,44,46,47,49,58,59 [30] COELOSIA Winnertz, 1863 flava (Staeger, 1840) - Bromeswell, 10 June 1990, P.J. Chandler TM 34 [1] fusca Bezzi, 1892 - Cavenham Heath, 30 May 1994, I. Perry TL 77,78,87 [1] ECTREPESTHONEURA Enderlein, 1911 colyeri Chandler, 1980 - The King’s Forest, 16 June 1991, I. Perry TL 87,88 [10] hirta (Winnertz, 1846) - Bromeswell, 10 June 1990, P.J. Chandler TM 34 [1] GRZEGORZEKIA Edwards, 1941 collaris (Meigen, 1818) - Bradfield Woods, 13 June 2007, I. Perry TL 95 [1] PALAEODOCOSIA Meunier, 1904 vittata (Coquillett, 1901) - Bradfield Woods, 9 October 1990, I. Perry TL 95 [1] SAIGUSAIA Vockeroth, 1980 flaviventris (Strobl, 1894) - Newbourne, 10 June 1990, P.J. Chandler TL 74,95 TM 24 [6] SYNAPHA Meigen, 1818 fasciata Meigen, 1818 - Walberswick, June 1988, A. Foster & D. Proctor TL 87,95,97 TM 24,34,47 [7] vitripennis (Meigen, 1818) - Barton Mills, 2 July 1954, J.E. Collin TL 77,78,86,87,95 TM 24,34,35,46,47 [33]

TETRAGONEURA Winnertz, 1846 sylvatica (Curtis, 1837) - Boyton, 10 July 1905, G.H. Verrall TL 77,78,87,88,95 TM 17,24,34,35,44,46,47,59 [28] LEIINAE DOCOSIA Winnertz, 1863 carbonaria Edwards, 1941 - Barton Mills, 9 May 1938, Verrall Collin collection TL 77,78,87 [8] flavicoxa Strobl, 1900 - Flatford, 29 August 1953, L. Parmenter TM 03 [1] fuscipes (von Roser, 1840) - West Stow, 20 April 2002, I. Perry TL 77[1] gilvipes (Haliday in Walker, 1856) - Walberswick, 11 July 1988, A. Foster & D. Procter TL 78,87 TM 47 [5] moravica Landrock, 1916 - Ampton, 1925, C.G. Nurse TL 78,87 [3] sciarina (Meigen, 1830) - Bentley Woods, 1901, C. Morley TL 78,86,95 TM 13 [10] Trans. Suffolk Nat. Soc. 55 (2019)

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GREENOMYIA Brunetti, 1912 mongolica Laštovka & Matile, 1974 - The King’s Forest, 13 July 2009, I. Perry TL 87 [1] LEIA Meigen, 1818 bilineata (Winnertz, 1863) - Walberswick, 1958, E.C.M. d’Assis-Fonseca TM 47 [1] bimaculata (Meigen, 1804) - Orford, July-August 1908, J.J.F.X. King TL 95 TM 26,44 [7] crucigera Zetterstedt, 1838 - Bradfield Woods, 19 June 2006, I. Perry TL 95 [2] earlier records in TSNS without details cylindrica (Winnertz, 1863) - Staverton Thicks, Wantisden, 22 August 1918, C. Morley TL 87 TM 25 [2] fascipennis Meigen, 1818 - Orford, July-August 1908, J.J.F.X. King TM 44,48 [2] longiseta Barendrecht, 1938 - Walberswick, August 1988, A. Foster & D. Procter TM 46,47 [7] subfasciata (Meigen, 1818) - Bradfield Woods, 19 June 2007, I. Perry TL 95 [1] earlier records in TSNS without details winthemii Lehmann, 1822 - Monk Soham, 4 June 1909, C. Morley TL 66,87,95 TM 26,46 [6] MEGOPHTHALMIDIA Dziedzicki, 1889 crassicornis (Curtis, 1837) - Harkstead, 1 September 1901, C. Morley TL 86 TM 13 [2] no recent records MYCETOPHILINAE Exechiini ALLODIA Winnertz, 1863 Subgenus ALLODIA sensu stricto anglofennica Edwards, 1921 - The King’s Forest, 4 November 2002, I. Perry TL 87 [1] embla Hackman, 1971 - Walberswick, August 1989, A. Foster & D. Procter TM 47 [2] lugens (Wiedemann, 1817) - Westleton, 19 September 1912, C. Morley TG 50 TL 77,78,86,87,95 TM 14,25,26,34,45,46,47 [40] lundstroemi Edwards, 1921 - The King’s Forest, 7 October 2002, I. Perry TL 87,95 [2] ornaticollis (Meigen, 1818) - Tuddenham, September 1914, J.E. Collin TL 66,77,78,86,87,95 TM 07,13,15,17,24,34,46,47,49 [49] truncata Edwards, 1921 - The King’s Forest, 20 April 2002, I. Perry TL 87 [3] earlier records in TSNS without details zaitzevi Kurina, 1998 - Mildenhall, 22 May 1909, J.W. Yerbury TL 77,95 TM 47 [3] Subgenus BRACHYCAMPTA Winnertz, 1863 alternans (Zetterstedt, 1838) - The King’s Forest, 4 November 2002, I. Perry TL 87 [1] earlier records in TSNS without details angulata (Lundström, 1913) - Walberswick, 1988-1989, A. Foster & D. Procter TM 47 [1] barbata (Lundström, 1909) - Tuddenham, July-August 1988, A. Foster & D. Procter TL 77,78 TM 47 [4] grata (Meigen, 1830) - Easton, August 1988, A. Foster & D. Procter TL 87,95 TM 17,25,47 [12]

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neglecta Edwards, 1925 - The King’s Forest, 18 July 2002, I. Perry TL 87 [1] protenta Laštovka & Matile, 1974 - Walberswick, August 1989, A. Foster & D. Procter TL 78 TM 47 [4] ALLODIOPSIS Tuomikoski, 1966 domestica (Meigen, 1830) - Belstead, September 1984, R.E. Evans TL 77,87 TM 14 [4] earlier records in TSNS without details rustica (Edwards, 1941) - Flatford, 13 July 1951, L. Parmenter TL 77,78,87,95 TM 03,04,35,46,47 [34] ANATELLA Winnertz, 1863 ciliata Winnertz, 1863 - Tuddenham, July-August 1988, A. Foster & D. Procter TL 77,78 [3] dampfi Landrock, 1924 - Walberswick, June 1988, A. Foster & D. Procter TM 47 [4] emergens Caspers, 1987 - Walberswick, 1988-1989, A. Foster & D. Procter TM 47 [1] flavomaculata Edwards, 1925 - Eriswell, 24 October 1934, J.E. Collin TL 77,87 [2] lenis Dziedzicki, 1923 - Cavenham Heath, 30 April 1994, I. Perry TL 77,87 [3] minuta (Staeger, 1840) - Tuddenham, July-August 1988, A. Foster & D. Procter TL 77 [1] pseudogibba Plassmann, 1977 - Tuddenham, June 1988, A. Foster & D. Procter TL 77,78 TM 47 [5] simpatica Dziedzicki, 1923 - The King’s Forest, 1 July 2000, I. Perry TL 87 [2] turi Dziedzicki, 1923 - West Stow, 1 November 2001, I. Perry TL 77,78 [2] unguigera Edwards, 1921 - Tuddenham, July-August 1988, A. Foster & D. Procter TL 77,78 TM 47 [6] BRACHYPEZA Winnertz, 1863 Subgenus BRACHYPEZA sensu stricto bisignata Winnertz, 1863 - The King’s Forest, 28 July 2002, I. Perry TL 87 [1] BREVICORNU Marshall, 1896 auriculatum (Edwards, 1925) - Walberswick, 1988-1989, A. Foster & D. Procter TL 87,97 TM 47 [5] fissicauda (Lundström, 1911) - Newmarket, 1894-1909, Verrall Collin collection TL 66,78,87,95 TM 47 57 [14] griseicolle (Staeger, 1840) - Tuddenham, 16 September 1891, Verrall Collin collection TL 66,77,78,86,87,95 TM 15,24,34,45,46,57 [48] intermedium (Santos Abreu, 1920) - The King’s Forest, 14 August 2002, I. Perry TL 87 [2] proximum (Staeger, 1840) - The King’s Forest, 28 July 2002, I. Perry TL 78,87 [3] sericoma (Meigen, 1830) - Mildenhall, 22 May 1909, J.W. Yerbury TL 77,78,87,95 TM 15,46,47,57 [40] verralli (Edwards, 1925) - Boyton, 27 July 1908, J.E. Collin TL 77,87,95 TM 34 [10] CORDYLA Meigen, 1803 brevicornis (Staeger, 1840) - The King’s Forest, 1 November 2004, I. Perry TL 78,87,95 [7] earlier records in TSNS without details crassicornis Meigen, 1818 - Ipswich, 10 June 1990, P.J. Chandler TL 77,78,86,87,95 TM 15,24,46,47 [27] earlier records in TSNS without details Trans. Suffolk Nat. Soc. 55 (2019)

Suffolk Natural History, Vol. 55 fissa Edwards, 1925 - Barnham, 22 June 1996, I. Perry TL 87,78,95 [3] flaviceps (Staeger, 1840) - Tuddenham, 1891, G.H.Verrall TL 77,78,87,88 [14] fusca Meigen, 1804 - Brandon, 26 October 2004, I. Perry TL 78 [2] insons Laštovka & Matile, 1974 - The King’s Forest, 4 November 2004, I. Perry TL 87 [1] murina Winnertz, 1863 - Ampton, 1916, C.G. Nurse TL 77,78,87,95 TM 34,47 [16] pusilla Edwards, 1925 - West Stow, 4 May 2002, I. Perry TL 77,87 [2] semiflava (Staeger, 1840) - Walberswick, 19 April 2018, I. Perry TM 47[1]

EXECHIA Winnertz, 1863 bicincta (Staeger, 1840) - Newmarket, 1888-1912, G.H. Verrall TL 66, 86 TM 46 [3] borealis Lundström, 1912 - Walberswick, 26 May 1973, A.M. Hutson TM 47 [1] cincta Winnertz, 1863 - Scotland Fen, Boyton, 19 October 1996, P.J. Chandler & M. Edwards TL 77,78,87,95 TM 34 [11] contaminata Winnertz, 1863 - Eastbridge, 18 October 1975, A.G. Irwin TL 77,78,87,95 TM 46 [11] dizona Edwards, 1924 - West Stow, 1 November 2001, I. Perry TL 77,78,87 [7] dorsalis (Staeger, 1840) - Herringfleet Woods, 3 April 1936, C. Morley TL 77,78,86,87,88,95 TM 34,35,45,49 [40] exigua Lundström, 1909 - Eriswell, 24 October 1934, J.E. Collin TL 77,78,87 [3] festiva Winnertz, 1863 - Scotland Fen, Boyton, 19 October 1996, P.J. Chandler & M. Edwards TL 77,78,87,88 [26] fusca (Meigen, 1804) - Washbrook, 27 March 1897, C. Morley TG 50 TL 66,77,78,86,87,88,95 TM 13,14,15,24,34,35,46,47 [67] lucidula (Zetterstedt, 1838) - Newmarket, 1888-1912, G.H. Verrall TL 66 [1] no recent records nigroscutellata Landrock, 1912 - Walberswick, 1988-1989, A. Foster & D. Procter TL 77,78,87 TM 47 [23] parva Lundström, 1909 - Tuddenham, 9 September 1915, C. Morley TL 78,87 [8] parvula (Zetterstedt, 1852) - Wangford Carr, July-August 1988, A. Foster & D. Procter TL 77,78,87 TM 49 [14] pseudocincta Strobl, 1910 - Newmarket, 23 September 1888, G.H. Verrall TL 66,78,87 [4] pseudofestiva Lackschewitz, 1937 - Tuddenham, July-August 1988, A. Foster & D. Procter TL 77,78 TM 47 [7] repanda Johannsen, 1912 - Walberswick, August 1988, A. Foster & D. Procter TL 77,87 TM 46,47 [12] repandoides Caspers, 1984 - Easton Broad, Covehithe, August 1982, P. Skidmore TL 86,87,95 TM 46,47,57 [7] separata Lundström, 1912 - Ampton, 1914, C.G. Nurse TL 77,78,87 [8] seriata (Meigen, 1830) - West Stow, 1 November 2001, I. Perry TL 77,78,87,95 [17] spinuligera Lundström, 1912 - Tuddenham, July-August 1988, A. Foster & D. Procter TL 77,78,87 TM 15,47 [9] earlier records in TSNS without details

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SUFFOLK DIPTERA CHECKLIST

EXECHIOPSIS Tuomikoski, 1966 Subgenus EXECHIOPSIS sensu stricto clypeata (Lundstrรถm, 1911) - Bradfield Woods, 26 October 2005, I. Perry TL 95 [2] fimbriata (Lundstrรถm, 1909) - Eriswell, 24 October 1934, J.E. Collin TL 77 [1] hammi (Edwards, 1925) - Tuddenham, July-August 1988, A. Foster & D. Procter TL 77,78, TM 47 [4] indecisa (Walker, 1856) - Tuddenham, 16 September 1891, G.H.Verrall TL 77 [1] no recent records intersecta (Meigen, 1818) - Minsmere, 18 October 1996, P.J. Chandler, R. Crossley, M. Edwards & K. Merrifield TL 87,95 TM 35,45,46 [5] earlier records in TSNS without specific details subulata (Winnertz, 1863) - Tunstall Forest, 19 October 1996, P.J. Chandler, K. Merrifield & R. Thomas TL78,87 TM 45 [5] Subgenus XENEXECHIA Tuomikoski, 1966 crucigera (Lundstrรถm, 1909) - Monk Soham, 30 October 1917, C. Morley TL 77,87 TM 26,46,47,59 [9] davatchii (Matile, 1969) - Brandon, 24 October 2010, I. Perry TL 78 [1] leptura (Meigen, 1830) - Newmarket, 1888-1912, G.H. Verrall TL 66,78 TM 34 [6] pollicata (Edwards, 1925) - Timworth, 19 February 1916, C.G. Nurse TL 86 [1] no recent records seducta (Plassmann, 1976) - Elveden, 20 September 2008, P.J. Chandler TL 78 [4] MYROSIA Tuomikoski, 1966 maculosa (Meigen, 1818) - Newmarket, 1888-1912, G.H. Verrall TL 66 [1] no recent records PSEUDEXECHIA Tuomikoski, 1966 parallela (Edwards, 1925) - Newmarket, 23 September 1888, G.H. Verrall TL 66,78 TM 47 [12] trisignata (Edwards, 1913) - Walberswick, August 1988, A. Foster & D. Proctor TL 77,78,87 TM 47 [5] trivittata (Staeger, 1840) - Blythburgh, 14 September 1912, C. Morley TL 66,77,78,86,87,95 TM 44,47,48 [15] tuomikoskii Kjรฆrandsen, 2009 - Bradfield Woods, 4 October 2006, I. Perry TL 78,95 [5] PSEUDOBRACHYPEZA Tuomikoski, 1966 helvetica (Walker, 1856) - Eastbridge, 18 October 1975, A.G. Irwin TL 87,95 TM 04,14,34,46,49 [22] RYMOSIA Winnertz, 1863 bifida Edwards, 1925 - Bradfield Woods, 17 October 2005, I. Perry TL 95 [4] fasciata (Meigen, 1804) - Newmarket, 10 February 1888, G.H. Verrall TG 50 TL 66,77,78,86,87,88,95 TM 37,45 [29] placida Winnertz, 1863 - North Cove, 17 October 1996, P.J. Chandler & M. Edwards TL 77,78,87,95 TM 49 [5]

Trans. Suffolk Nat. Soc. 55 (2019)

Suffolk Natural History, Vol. 55 signatipes (van der Wulp, 1859) - Scotland Fen, Boyton, 19 October 1996, P.J. Chandler & M. Edwards TL 87,88,95 TM 34 [9] spinipes Winnertz, 1863 - Ickworth, 8 October 2004, I. Perry TL 78,86,87,95 [10] virens Dziedzicki, 1910 - Bradfield Woods, 4 October 2006, I. Perry TL 95 [1]

STIGMATOMERIA Tuomikoski,1966) crassicornis (Stannius, 1831) - Staverton Thicks, Wantisden, 1907-1911, Verrall Collin collection TL 65,66,77,78,86,87,95 TM 03,26,35,46,47 [44] SYNPLASTA Skuse, 1890 gracilis (Winnertz, 1863) - Dunwich Forest, 26 May 1974, A.M. Hutson TL 77,78,87 TM 24,35,46,47,49 [14] ingeniosa (Kidd, 1969) - Wangford Carr, Brandon, June-September 1988, A. Foster & D. Proctor TL 78,95 TM 45 [3] TARNANIA Tuomikoski, 1966 fenestralis (Meigen, 1818) - Monk Soham, 26 February 1935, C. Morley TG 50 TL 77,78,86,87 TM 14,26,47 [14] nemoralis (Edwards, 1941) - Minsmere, 18 October 1996, P.J. Chandler, R. Crossley, M. Edwards & K. Merrifield TL 78,87 TM 46 [3] Mycetophilini DYNATOSOMA Winnertz, 1863 cochleare Strobl, 1895 - The King’s Forest, 25 September 2002, I. Perry TL 87 [3] fuscicorne (Meigen, 1818) - Flatford, 17 July 1951, L. Parmenter TL 77,78,87,95 TM 03,04,35,44,47 [18] reciprocum (Walker, 1848) - The King’s Forest, 21 June 1998, I. Perry TL 78,87 TM 04 [6] EPICYPTA Winnertz, 1863 aterrima (Zetterstedt, 1852) - Scotland Fen, Boyton,19 October 1996, P.J. Chandler & M. Edwards TL 78,86,87,95 TM 34 [10] limnophila Chandler, 1981 - Walberswick, June-August 1988, A. Foster & D. Proctor TL 77 TM 47 [10] MYCETOPHILA Meigen, 1803 abiecta (Laštovka, 1963) - The King’s Forest, 29 October 2001, I. Perry TL 87 [5] adumbrata Mik, 1884 - The King’s Forest, 6 August 1994, I. Perry TL 87 TM 34 [17] alea Laffoon, 1965 - Tuddenham, 16 September 1891, Verrall Collin collection TL 66,67,78,95 TM 46 [8] autumnalis Lundström, 1909 - The King’s Forest, 28 October 2001, I. Perry TL 78,87 [10] blanda Winnertz, 1863 - Eriswell, 24 October 1934, J.E. Collin TL 77,78,87 [7] britannica Laštovka & Kidd, 1975 - Ampton, 1913-1914, C.G. Nurse TL 66,77,78,86,87,88,95 TM 04,15,46 [40] caudata Staeger, 1840 - The King’s Forest, 2 August 1987, I. Perry TL 78,87 TM 34 [13] cingulum Meigen, 1830 - Bentley Woods, April 1897, C. Morley TL 88,86,87,95 TM 03,04,07,15,25,44 [19]

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confusa Dziedzicki, 1884 - Eriswell, 24 October 1934, J.E. Collin TL 77 TM 24,26 [4] curviseta Lundström, 1911 - Ampton, 1916, C.G. Nurse TL 77,78,86,87,88,95 TM 04,45,46,49 [38] deflexa Chandler, 2001 - Brandon, 18 April 2011, I. Perry TL 78 [1] dentata Lundström, 1913 - The King’s Forest, 25 September 2002, I. Perry TL 78,87,95 TM 35,46 [14] dziedzickii Chandler, 1977 - Tuddenham, 16 September 1891, Verrall Collin collection TL 66,77,78,87 TM 34,46 edwardsi Lundström, 1913 - Tuddenham Fen, 20 October 1990, P.J. Chandler & I.F.G. McLean TL 77,86,87,95 TM 04,34 [24] eppingensis Chandler, 2001- The King’s Forest, 19 September 2002, I. Perry TL 87,95 [6] forcipata Lundström, 1913 - Barton Mills, 1910-1932, J.E. Collin TL 77,78,87,95 TM 04,24 [20] formosa Lundström, 1911 - The King’s Forest, 16 June 1991, I. Perry TL 78,86,87,95 TM 04,14 [23] fraterna Winnertz, 1863 - Herringfleet, 3 April 1936, C. Morley TL 77,78,87,95 TM 04,24,34,35,45,49,58,59 [35] fungorum (De Geer, 1776) - Tuddenham, 16 September 1891, Verrall Collin collection TL 66,77,78,87,95 TM 15,17,24,34,35,44,47,49,57,58 [51] gibbula Edwards, 1925 - The King’s Forest, 13 September 2002, I. Perry TL 87,95 [7] ichneumonea Say, 1823 - West Stow, 19 July 1918, C.G. Nurse TL 77,78,87,95 TM 14,44,46,47,49 [13] lastovkai Caspers, 1984 - Staverton Thicks, Wantisden, 6 June 1910, Verrall Collin collection TL 78, TM 35 [2] luctuosa Meigen, 1830 - Timworth, 1916, C.G. Nurse TL 77,78,86,87,88,95 TM 15,46,58 [23] lunata Meigen, 1804 - Boyton, 27 August 1907, J.E. Collin TL 78,86,87,95 TM 34 [18] marginata Winnertz, 1863 - Timworth, 1916, C.G. Nurse TL 77,78,86,95 TM 24,35,46 [33] mitis (Johannsen, 1912) - Shrubland Park, Barham, 20 October 1996, I.F.G. McLean TL 87 TM 15 [4] occultans Lundström, 1913 - Ampton, 1916, C.G. Nurse TL 87 [2] ocellus Walker, 1848 - Newmarket, 5 February 1887, Verrall Collin collection TL 65,66,77,78,86,87,95 TM 04,15,24,26,34,45,46,47,48 [52] ornata Stephens, 1846 - Flatford, 17 May 1951, L. Parmenter TL 87,95 TM 03,04,15,34,58 [9] perpallida Chandler, 1993 - Tuddenham, 16 September 1891, Verrall Collin collection TL 77,78,88,95 [7] pictula Meigen, 1830 - Newmarket, 1888-1912, G.H. Verrall TL 66,77,87,95 [14] pumila Winnertz, 1863 - Tuddenham Fen, 20 October 1990, P.J. Chandler & I.F.G. McLean TL 77,95 [5] rudis Winnertz, 1863 - Scotland Fen, Boyton, 19 October 1996, P. J. Chandler & M. Edwards TL 88 TM 34 [6] Trans. Suffolk Nat. Soc. 55 (2019)

Suffolk Natural History, Vol. 55 ruficollis Meigen, 1818 - Boyton, 27 August 1908, Verrall Collin collection TL 77,87 TM 34 [4] scotica Edwards, 1941 - Bradfield Woods, 26 October 2005, I. Perry TL 95 [2] sepulta (Laffoon, 1957) - The King’s Forest, 28 October 2001, I. Perry TL 87 [2] signatoides Dziedzicki, 1884 - Walberswick, 15 June 1989, A. Foster & D. Proctor TL 78,87,88,95 TM 47 [16] sordida van der Wulp, 1874 - Eastbridge, 18 October 1975, A.G. Irwin TL 78,95 TM 46,47 [6] stolida Walker, 1856 - Ampton, 1914, C.G. Nurse TL 87 [1] no recent records stricklandi (Laffoon, 1957) - West Stow, 20 April 2002, I. Perry TL 77,87 [6] strigata Staeger, 1840 - Mildenhall, 22 May 1909, J.W. Yerbury TL 77,87,95 TM 26 [7] stylata (Dziedzicki, 1884) - Ickworth, 19 September 2005, I. Perry TL 86 [1] sublunata Zaitzev, 1998 - Brandon, 18 April 2011, I. Perry TL 78 [1] sumavica (Laštovka, 1963) - Staverton Thicks, Boyton, 19 October 1996, P.J. Chandler, C. McLean & I.F.G. McLean TL 88 TM 35 [2] tridentata Lundström, 1911 - Brandeston, 16 September 1972, P.J. Chandler TL 86,87,95 TM 26 [13] trinotata Staeger, 1840 - Cavenham Heath, 30 April 1994, I. Perry TL 77,78,86,87,95 TM 34,47 [14] unicolor Stannius, 1831 - Tuddenham Fen, 20 October 1990, P.J. Chandler & I.F.G. McLean TL 77,78.87,95 TM 34,47,49,59 [16] unipunctata Meigen, 1818 - Walberswick, 26 May 1973, A.M. Hutson TL 78,87 TM 34,45,46,47,49 [14] vittipes Zetterstedt, 1852 - The King’s Forest, 27 July 1991, I. Perry TL 87 [1]

PHRONIA Winnertz, 1863 basalis Winnertz, 1863 - Staverton Thicks, Boyton, 2 February 1910, Verrall Collin collection TL 77,78,87,95 TM 35,58 [13] biarcuata (Becker, 1908) - Timworth, 1916, C.G. Nurse TL 77,78,86,87,95 TM 04,34,35,45,46,47 [40] braueri Dziedzicki, 1889 - Walberswick, 26 May 1973, A.M. Hutson TL 78,87.95 TM 35,47 [7] cinerascens Winnertz, 1863 - Brookhill Wood, Ipswich, 10 June 1990, P.J. Chandler TL 77,78,87,88 TM 24,35 [20] conformis (Walker, 1856) - Newmarket, September 1888, G.H. Verrall TL 66,77,78,86,87,95 TM 24,35,46,47,48 [32] coritanica Chandler, 1992 - Staverton Thicks, Boyton, 25 April 1952, Verrall Collin collection TL 77,87,95 TM 35 [13] disgrega Dziedzicki, 1889 - The King’s Forest, 1 July 2000, I. Perry TL 77,87 [2] exigua (Zetterstedt, 1852) - Newmarket, September 1888, G.H. Verrall TL 66,78,87 [4] flavipes Winnertz, 1863 - The King’s Forest, 25 May 2002, I. Perry TL 87 [1] forcipata Winnertz, 1863 - Barton Mills, 3 April 1994, I.F.G. McLean TL 77,87,95 TM 34 [15] humeralis Winnertz, 1863 - Mildenhall, 22 May 1905, J.W. Yerbury TL 77,78,87,95 TM 24,34,45,46 [26] Trans. Suffolk Nat. Soc. 55 (2019)

SUFFOLK DIPTERA CHECKLIST

nigricornis (Zetterstedt, 1852) - Staverton Thicks, Boyton, 15 April 1952, Verrall Collin collection TL 77,87 TM 15,34,35,47,77,87 [18] nitidiventris (van der Wulp, 1859) - Cavenham Heath, 14 May 1994, I. Perry TL 77,78,87,95 [19] notata Dziedzicki, 1889 - Coddenham, August 1992. P. Skidmore TL 78,87,95 TM 15,35 [7] obtusa Winnertz, 1863 - Cavenham Heath, 30 April 1994, I. Perry TL 77,87 [6] earlier records in TSNS without details portschinskyi Dziedzicki, 1889 - Walberswick, 1 June 1988, A. Foster & D. Proctor TM 46,47 [3] siebeckii Dziedzicki, 1889 - The King’s Forest, 30 April 1994, I. Perry TL 87 [3] strenua Winnertz, 1863 - The King’s Forest, 4 June 2000, I. Perry TL 86,87,88,95 [9] tenuis Winnertz, 1863 - Cavenham Heath, 30 April 1994, I. Perry TL 77,78 [4] triangularis Winnertz, 1863 - The King’s Forest, 16 June 1991, I. Perry TL 77,78,87 [11] PLATUROCYPTA Enderlein, 1910 punctum (Stannius, 1831) - Orford, July - August 1908, J.J.F.X. King TL78,87,95 TM 44 [14] testata (Edwards, 1924) - Tuddenham, June - August 1988, A. Foster & D. Proctor TL 77,78,87,95 TM 24,35 [24] SCEPTONIA Winnertz, 1863 concolor Winnertz, 1863 - Mildenhall, (no date available), G.H.Verrall TL 77 [1] no recent records costata (van der Wulp, 1859) - The King’s Forest, 16 June 1991, I. Perry TL 87,88 [2] cryptocauda Chandler, 1991 - Lavenham, 17 July 2003, P.J. Chandler TL 78,84 [3] flavipuncta Edwards, 1925 - Mildenhall, 24 June 1909, J.W. Yerbury TL 77 [1] no recent records fumipes Edwards, 1925 - Cavenham Heath, 30 April 1994, I. Perry TL 77,78,87 TM 45 [7] fuscipalpis Edwards, 1925 - Walberswick, 26 May 1973, A.M. Hutson TM 47 [1] membranacea Edwards, 1925 - The King’s Forest, 27 July 1991, I. Perry TL 78,87,95 TM 34,35,46,59 [14] nigra (Meigen, 1804) - Timworth, 1915, C.G. Nurse TL 77,78,86,87 TM 24,34,46 [19] TRICHONTA Winnertz, 1863 apicalis Strobl, 1898 - Brandon, 1 May 1994, I. Perry TL 78 [1] atricauda (Zetterstedt, 1852) - Monk Soham, July 1916, C. Morley TL 77,78,87 TM 26,47 [12] bicolor Landrock, 1912 - The King’s Forest, 30 April 1994, I. Perry TL 87 [1] earlier records in TSNS without details clavigera Lundström, 1913 - The King’s Forest, 30 April 1994, I. Perry TL 87 [1] falcata Lundström, 1911 - Minsmere, 18 October 1996, I.F.G. McLean & A.E. Stubbs TL 77 TM 46 [2] foeda Loew, 1869 - Bentley Woods, 5 March 1935, C. Morley TM 13 [1] no recent records Trans. Suffolk Nat. Soc. 55 (2019)

Suffolk Natural History, Vol. 55 fragilis Gagné, 1981 - Cavenham Heath, 30 April 1994, I. Perry TL 77,78,87 [3] melanura (Staeger, 1840) - Staverton Thicks, Boyton, 19 October 1996, P.J. Chandler, C. McLean & I.F.G. McLean TL 77,78,87,88 TM 35 [16] nigritula Edwards, 1925 - Walberswick, August 1989, A. Foster & D. Proctor TM 47 [2] submaculata (Staeger, 1840) - Southwold, 19 September 1919, C. Morley TL 77,87,95 TM 57 [7] terminalis (Walker, 1856) - Cavenham Heath, 30 April 1994, I. Perry TL 77,87,95 [6] vitta (Meigen, 1830) - Cavenham Heath, 30 April 1994, I. Perry TL 77,78,87,95 [14]

ZYGOMYIA Winnertz, 1863 humeralis (Wiedemann, 1817) - Barnham, 14 August 1970, H.H. Burns & Huxley TL 77,78,87,95 TM 15,35,47 [34] kiddi Chandler, 1991 - Brandon, 19 September 2011, I. Perry TL 78 [1] notata (Stannius, 1831) - Lakenheath, June-September 1988, A. Foster & D. Proctor TL 77,78,87 TM 47 [7] pictipennis (Staeger, 1840) - Bentley Woods, February 1899, C. Morley TL 77,78,87,88 TM 34,45,46,47,49 [19] pseudohumeralis Caspers, 1980 - Wangford Carr, Brandon, June-September 1988, A. Foster D &. Proctor TL 77,78 TM 15,47 [7] valida Winnertz, 1863 - Newmarket, 1886-1907, Verrall Collin collection TL 66,77,78,86,87,95 TM 04,15,34,45,46,47,59 [24] vara (Staeger, 1840) - Newmarket, 6 December 1886, Verrall Collin collection TL 66,77,78,87,95 TM 34,35,44,45,46,59 [21] MYCOMYINAE MYCOMYA Rondani, 1856 Subgenus CYMOMYA Väisänen, 1984 circumdata (Staeger, 1840) - Staverton Thicks, Boyton, 7 September 1916, C. Morley TL 77,78,87,95 TM 35,48 [16] Subgenus MYCOMYA sensu stricto annulata (Meigen, 1818) - Benacre, 28 May 1973, A.M. Hutson TL 77,95 TM 46,47,49,59 [18] cinerascens (Macquart, 1826) - Ampton, 1916-1921, C.G. Nurse TL77,78,87,95,97 TM 03,24,34,35,45,46,47,48 [52] flavicollis (Zetterstedt, 1852 - Bradfield Woods, 11 October 2005, I. Perry TL 86,95 [7] marginata (Meigen, 1818) - Tuddenham, 16 September 1891, Verrall Collin collection TL 77,78,87,95 TM 34,37,45,46,47 [34] neohyalinata Väisänen, 1984 - Minsmere, 18 October 1996, P.J. Chandler, R. Crossley, M. Edwards & K. Merrifield TL 77,78 TM 34,45,46 [9] nitida (Zetterstedt, 1852) - Bradfield Woods, 23 May 1991, I. Perry TL 95 [1] parva (Dziedzicki, 1885 - The King’s Forest, 30 April 1994, I. Perry TL 77,78,87,89,95 [8] prominens (Lundström, 1913) - The King’s Forest, 25 September 2002, I. Perry TL 87,95 [3] punctata (Meigen, 1804) - The King’s Forest, 25 September 2002, I. Perry TL 87,95 [3]

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sigma Johannsen, 1910 - The King’s Forest, 10 October 2002, I. Perry TL 78,87,95 [3] tenuis (Walker, 1856) - The King’s Forest, 4 May 2002, I. Perry TL 78,86,87,95 [10] tumida (Winnertz, 1863) - The King’s Forest, 29 October 2001, I. Perry TL 87,95 TM 47 [4] wankowiczii (Dziedzicki, 1885) - Bradfield Woods, 11 July 1990, B.R. Laurence TL 78,87,95 TM 46 [5] winnertzi (Dziedzicki, 1885) - Thornham Magna, 25 April 1991, P. Withers TL 77,78,86,87,95 [16] Subgenus MYCOMYOPSIS Väisänen, 1984 affinis (Staeger, 1840) - Walberswick, 1988-1989, A. Foster & D. Proctor TM 47 [1] maura (Walker, 1856) - Tuddenham Heath, 15 April 1981, I. Perry TL 87,95 TM 47 [14] trilineata (Zetterstedt, 1838 - Bradfield Woods, 19 June 2007, I. Perry TL 95 [1] NEOEMPHERIA Osten Sacken, 1878 pictipennis (Haliday, 1833) - Scotland Fen, Boyton, 19 October 1996, P.J. Chandler & M. Edwards TL 78,86,87 TM 34 [8] SCIOPHILINAE ACNEMIA Winnertz, 1863 nitidicollis (Meigen, 1818) - Timworth, 1916, C.G. Nurse TL 77,78,86,87,88,95 TM 24,34,35,44,47 [37]

ALLOCOTOCERA Mik, 1886 pulchella (Curtis, 1837) - Barton Mills, 24 June 1960, E.C.M. d’Assis-Fonseca TL 77,95 [2] AZANA Walker, 1856 Subgenus AZANA sensu stricto anomala (Staeger, 1840) - Newmarket, 1888-1912, G.H. Verrall TL 66 [1] no recent records COELOPHTHINIA Edwards, 1941 thoracica (Winnertz, 1863) - The King’s Forest, 9 October 1994, I. Perry TL 78,87,95 TM 45,46 [16] LEPTOMORPHUS Curtis, 1831 Subgenus LEPTOMORPHUS sensu stricto walkeri Curtis, 1831 - Freston, 22 July 1904, C. Morley TM 13,26 [2] no recent records MEGALOPELMA Enderlein, 1911 nigroclavatum (Strobl, 1910) - Brookhill Woods, Ipswich, 10 June 1990, P.J. Chandler TL 77,78 TM 24 [4] MONOCLONA Mik, 1886 rufilatera (Walker, 1837) - Orford, July-August 1908, J.J.F.X. King TL 77,78,86,87,89,95 TM 04,44,47 [17]

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NEURATELIA Rondani, 1856 nemoralis (Meigen, 1818) - Bentley Woods, 29 May 1902, C. Morley TL 95 TM 13 [2] PHTHINIA Winnertz, 1863 mira (Ostroverkhova, 1977) - Ickworth, 1 October 2004, I. Perry TL 78,86 [4] winnertzi Mik, 1869 - Oulton, 17 October 1996, P.J. Chandler TL 77,87 TM 04,59 [4] SCIOPHILA Meigen, 1818 fenestella Curtis, 1837 - Cavenham Heath, 30 April 1994, I. Perry TL 77,95 [2] geniculata Zetterstedt, 1838 - Barking, 4 November 1945, C. Morley TM 05,26 [2] no recent records hirta Meigen, 1818 - Newmarket, May 1887, G.H. Verrall TL 66,95 [2] interrupta (Winnertz, 1863) - Mildenhall, 13 May1909, J.W. Yerbury TL 77 [1] no recent records lutea Macquart, 1826 - Walberswick, June-August 1988, A. Foster & D. Proctor TL 87,95 TM 47 [4] earlier records in TSNS without details nigronitida Landrock, 1925 - Aldeburgh, July 1907, J.W. Yerbury TL 77,87 TM 45 [3] parviareolata Santos Abreu, 1920 - Newmarket, 8 May 1886, Verrall Collin collection TL 66 [5] no recent records pomacea Chandler, 2006 - Holton, 1992, M. Ellis & P. Ellis TM 47 [1] thoracica Staeger, 1840 - Barton Mills, 27 July 1989, P.J. Chandler TL 77 [1] varia (Winnertz, 1863 - Monk Soham, 1940, C. Morley TM 26 [1] no recent records

SCIARIDAE BRADYSIA Winnertz, 1867 aprica (Winnertz, 1867) - Bradfield Woods, 1977-1984, B.R. Laurence TL 95 TM 07 [2] cinerascens (Grzegorzek, 1884) -Tunstall Common, 19 October 1996, P.J. Chandler TM 34,35 [3] confinis (Winnertz, 1867) - Newmarket, 29 April 1889, G.H. Verrall TL 66,95 [2] fungicola (Winnertz, 1867) - Redgrave, 24 May 1987, P. Withers TM 07 [1] giraudii (Egger, 1862) - Flatford, 12 July 1981, no collector details TM 03 [1] nitidicollis (Meigen, 1818) - Kessingland, April 1933, Edwards TL 95 TM 07,58 [8] nocturna Tuomikoski, 1960 - Scotland Fens, Boyton, 10 November 1996, P.J. Chandler TM 34 [1] normalis Frey, 1948 - Walberswick, June 1988, A. Foster & D. Procter TM 47 [1] pallipes (Fabricius, 1787) -Thornham Magna, 23 February 1990, P. Withers TM 17 [1] pectoralis (Staeger, 1840) - Santon Downham, August 1935, F.W. Edwards TL 88 [1] placida (Winnertz, 1867) - Walberswick, June 1988, A. Foster & D. Procter TL 97 TM 07,17,46,47 [6] polonica (Lengersdorf, 1929) - Thornham Magna, 20 May 1989, P. Withers TM 17 [1] praecox (Meigen, 1818) - Minsmere, 2004, D. Gibbs TM 46 [1] rectinervis Frey, 1948 - Bradfield Woods, October 1990, B.R. Laurence TL 95,97 [2] scabricornis Tuomikoski, 1960 - Walberswick, 15 June 1988, A. Foster & D. Procter TM 47[1] tilicola (Loew, 1850) - Market Weston, 6 September 1988, R.E. Evans TL 97 TM 17 [1]

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trivittata (Staeger, 1840) - Orford, 19 July 1908, J.J.F.X. King TL 78,95 TM 49 [3 vagans (Winnertz, 1868) - Occold, 10 May 1990, P. Withers TL 95 TM 17 [5] BRADYSIOPSIS Tuomikoski, 1960 vittata (Meigen, 1830) - Bradfield Woods, July 1990, B.R. Laurence TL 95 [2] CORYNOPTERA Winnertz, 1867 Subgenus CORYNOPTERA sensu stricto cuniculata (Lengersdorf, 1942) - Monk Soham, 5 June 1915, C. Morley TM 26 [1] no recent records curvispinosa Freeman, 1983 - Santon Downham, 15 August 1970, A.M. Hutson TL 88 [1] flavicauda (Zetterstedt, 1855) - Bradfield Woods, May 1991, B.R. Laurence TL 95 [1] flavosignata Menzel & Heller in Menzel, Smith & Chandler, 2006 - Bradfield Woods, July 1990, B.R. Laurence TL 95 [1] inundata Fritz, 1982 - Bradfield Woods, July 1990, B.R. Laurence TL 77,95 [2] obscuripila Tuomikoski, 1960 - Lakenheath, September 1988, A. Foster & D. Procter TL 78 [1] perpusilla Winnertz, 1867 - Lakenheath, September 1988, A. Foster & D. Procter TL 78 [1] saccata Tuomikoski, 1960 - Bradfield Woods, 16 July 1990, B.R. Laurence TL 95 [4] subfurcifera Mohrig & Hรถvemeyer, 1992 - Bradfield Woods, July 1990, B.R. Laurence TL 95 [1] trepida (Winnertz, 1867) - Santon Downham, 15 August 1970, A.M. Hutson TL 78,88 [2] CRATYNA Winnertz, 1867 Subgenus CRATYNA sensu stricto ambigua (Lengersdorf, 1934) - Bradfield Woods, May 1991, B.R. Laurence TL 95 [1] uliginosa (Lengersdorf, 1929) -Tuddenham Fen, 20 October 1990, P.J. Chandler TL 77 [1] wasmanni (Schmitz, 1919) - Bradfield Woods, July 1990, B.R. Laurence TL 95 [3] Subgenus SPATHOBDELLA Frey, 1948 falcifera (Lengersdorf, 1933) - Market Weston, 12 May 1991, P. Withers TL 97 [1] keilini (Edwards, 1915) - Barton Mills, 8 May 1915, D. Keilin TL 77, 95 [2] nobilis (Winnertz, 1867) - Reydon Wood, 11 May 1986, P. Withers TM 47 [1] CTENOSCIARA Tuomikoski, 1960 hyalipennis (Meigen, 1804) - Walberswick, 7 March 1973, S.J Chambers TM 47 [1] EPIDAPUS Haliday in Walker, 1851 Subgenus EPIDAPUS sensu stricto atomarius (De Geer, 1778) - Bradfield Woods, May 1991, B.R. Laurence TL 95 [3] gracilis (Walker, 1848) - Bradfield Woods, October 1990, B.R. Laurence TL 95 [1] microthorax (Bรถrner, 1903) - Bradfield Woods, April 1990, B.R. Laurence TL 95 [1] schillei (Bรถrner, 1903) - Bradfield Woods, May 1991, B.R. Laurence TL 95 [1]

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LEPTOSCIARELLA Tuomikoski, 1960 Subgenus HIRTIPENNIA Mohrig & Menzel, 1997 hirtipennis (Zetterstedt, 1838) - Ipswich, April 1895, C. Morley TM 14 [1] no recent records Subgenus LEPTOSCIARELLA sensu stricto nigrosetosa (Freeman, 1990) - Bradfield Woods, May 1991, B.R. Laurence TL 95 [1] rejecta (Winnertz, 1867) - Newmarket, 5 September 1894, G.H. Verrall TL 66 [1] no recent records trochanterata (Zetterstedt, 1851) - Parham, 5 May 1998, C. Hitch & P.M.E. Bennett TL 85 TM 36 [2] LYCORIELLA Frey, 1942 Subgenus HEMINEURINA Frey, 1942 conspicua (Winnertz, 1867) - Blaxhall Common, 7 August 1988, P. Kirby TM 35 [1] Subgenus LYCORIELLA sensu stricto castanescens (Lengersdorf, 1940) - Market Weston, 6 September 1988, P. Withers TL 95,97 TM 23 [5] ingenua (Dufour, 1839) - Belstead, 1984, R.E. Evans TM 14,35 [2] PHYTOSCIARA Frey, 1942 Subgenus DOLICHOSCIARA Tuomikoski, 1960 flavipes (Meigen, 1804) - Newmarket, 20 April 1901, G.H. Verrall TL 66 TM 35 [2] no recent records PSEUDOLYCORIELLA Menzel & Mohrig, 1998 subbruckii (Mohrig & Hรถvemeyer, 1992) - Walberswick, September 1988, A. Foster & D. Procter TM 46,47 [2] SCATOPSCIARA Edwards, 1927 Subgenus SCATOPSCIARA sensu stricto atomaria (Zetterstedt, 1851) - Thornham Magna, 14 March 1987, P. Withers TL 88,95 TM 17 [5] calamophila Frey, 1948 - Stallode Wash, Lakenheath, June-August 1988, A. Foster & D. Procter TL 78 [1] edwardsi Freeman, 1983 - Market Weston, 12 May 1991, P. Withers TL 97 [1] fluviatiliformis Mohrig & Mamaev, 1987 - Bradfield Woods, July 1990, B.R. Laurence TL 95 [1] nana (Winnertz, 1871) - Newmarket, 25 July 1893, G.H. Verrall TL 66,95 TM 07 [3] neglecta Menzel & Mohrig, 1998 - Bradfield Woods, August 1991, B.R. Laurence TL 95 [1] pusilla (Meigen, 1818) - Bradfield Woods, August 1988, B.R. Laurence TL 95 TM 03 [1] tricuspidata (Winnertz, 1867) - Bradfield Woods, 1977-1984, B.R. Laurence TL 95 [1] vitripennis (Meigen, 1818) - Wangford, 12 July 1973, Rogers, Fitton & Day TL 78 TM 17 [4]

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Subgenus XENOPYGINA Frey, 1948 fritzi Mohrig & Menzel, 1992 - Walberswick, June 1988, A. Foster & D. Procter TM 47 [1] simillima (Tuomikoski, 1960) - Walberswick, June 1988, A. Foster & D. Procter TM 47 [1] weiperti Menzel & Mohrig, 1991 - Lakenheath, September 1988, A. Foster & D. Procter TL 78 [1] SCHWENCKFELDINA Frey, 1942 carbonaria (Meigen, 1830) - Newmarket, 19 May 1884, G.H. Verrall TL 66,77,95 TM 15,17,26,46 [12] SCIARA Meigen, 1803 hemerobioides (Scopoli, 1763) - East Bergholt, July 1970, Lane TM 03,46,47 [5] earlier records in TSNS without details humeralis Zetterstedt, 1851- Walberswick, June 1988, A. Foster & D. Procter TM 46,47 [3] SCYTHROPOCHROA Enderlein, 1911 radialis Lengersdorf, 1926 - Santon Downham, 5 September 1970, no collector details TL 88 [1] TRICHOSIA Winnertz, 1867 Subgenus TRICHOSIA sensu stricto glabra (Meigen, 1830) - Ipswich, 10 May 1898, C. Morley TM 14,26 [2] no recent records morio (Fabricius, 1794) - Staverton Thicks, Wantisden, 1 June 1952, E.C.M. d’Assis-Fonseca TM 07,35,47 [3] splendens Winnertz, 1867 - Staverton Thicks, Wantisden, 1 June 1952, E.C.M. d’Assis-Fonseca TM 35 [1] ZYGONEURA Meigen, 1830 Subgenus ZYGONEURA sensu stricto sciarina Meigen, 1830 - Reydon Wood, 11 May 1996, P. Withers TM 34,47 [2]

CECIDOMYIIDAE Supertribe Cecidomyiidi Asphondyliini ASPHONDYLIA Loew, 1850 melanopus Kieffer, 1890 - Kelsale, 19 August 2015, J.P. Bowdey, Host: Lotus corniculatus TM 36 [1] sarothamni (Loew, 1850) - Ipswich, 21 July 2009, J.P. Bowdey, Host: Cytisus scoparius TM 14,45,46,47 [5] ulicis Trail, 1873 - Southwold, September 1939, C. Morley. Host: Ulex europaeus TM 45,46,59 [8] Schizomyiina KIEFFERIA Mik, 1895 pericarpiicola (Bremi, 1847) - Holbrook Bay, 1 August 2009, J.P. Bowdey. Host: Daucus carota TM 13,23,36,46 [4] Trans. Suffolk Nat. Soc. 55 (2019)

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SCHIZOMYIA Kieffer, 1889 galiorum Kieffer, 1889) - Sizewell, 18 July 2016, J.P. Bowdey Host: Galium verum TM 36,46 [2] Cecidomyiini ANISOSTEPHUS Rübsaamen, 1917 betulinus (Kieffer, 1889) - Bradfield Woods, 1990-1991, B.R. Laurence, Host: Betula spp. TL 95 [1] CONTARINIA Rondani, 1860 acerplicans (Kieffer, 1889) - Shottisham, 5 December 2018, R. Ruffell, Host: Acer pseudoplatanus TM 34 [1] artemisiae Rübsaamen, 1917 - Gunton Cliffs, 20 July 2014, J.P. Bowdey, Host: Artemisia vulgaris TL 98 TM 59 [2] barbichei (Kieffer, 1890) - Bradfield Woods, 20 September 2014, BPGS meeting, Host: Lotus uliginosus TL 95 [1] coryli (Kaltenbach, 1859) - Blundeston, April 1935, E.A. Ellis, Host: Corylus avellana TM 36,59 [1] craccae (Loew, 1850) - Kelsale, 18 July 2016, J.P. Bowdey, Host: Vicia cracca TM 36 [1] jacobaeae (Loew, 1850) - Belton, 15 July 1999, R. Hancy. Host: Senecio spp. TG 40 [1] loti (De Geer, 1776) - Alton Water, 8 July 2011, J.P. Bowdey, Host: Lotus corniculatus TM 13,36 [2] medicaginis Kieffer, 1895 - Kelsale, 10 August 2019, J.P. Bowdey, Host: Medicago sativa TM 46 [1] nicolayi (Rübsaamen, 1895) - Brantham, 14 June 2008, J.P. Bowdey, Host: Heracleum sphondylium TM 13,36 [3] pyrivora (Riley, 1886) - Saxmundham, 13 June 2017, J.P. Bowdey, Host: Pyrus domestica TM 36 [3] quinquenotata (F. Löw, 1888) - Ipswich, 10 July 1995, no collector details, Host: Hemerocallis fulva TL 86,87,96 TM 03,14,36,47 [15] steini (Karsch, 1881) - Knettishall Heath, 21 September 2014, BPGS meeting, Host: Silene dioica TL 98 TM 46 [2] tiliarum (Kieffer, 1890) - Minsmere, 15 June 1990, E. Beaumont, Host: Tilia sp. TM 04,15,36,46 [9] tritici (Kirby, 1798) - Barnham, May 1799, W. Kirby, Host: Triticum aestivum TM 15 [1] no recent records MACRODIPLOSIS Kieffer, 1895 pustularis (Bremi, 1847) - Iken Cliff, 9 August 1992, P. Lee, Host: Quercus sp. TG 40, TL 95 TM 03,04,07,13,15,24,34,35,46,49 [19] roboris (Hardy, 1854) - Bradfield Woods, 14 July 1994, R. Hancy, Host: Quercus sp. TG 40 TL 86,95 TM 03,13,14,15,24,34,36,46,47,49 [18] Clinodiplosini AMETRODIPLOSIS Rübsaamen, 1910 thalictricola (Rübsaamen, 1895) - Bungay, July 1932, E.A. Ellis, Host: Thalictrum flavum TM 38 [1] no recent records Trans. Suffolk Nat. Soc. 55 (2019)

SUFFOLK DIPTERA CHECKLIST

PARALLELODIPLOSIS Rübsaamen, 1910 galliperda (F. Löw, 1889) - Flatford, September 2008, BPGS meeting, Host: Neuroterus quercusbaccarum on Quercus TM 03 [1] Lopesiini OBOLODIPLOSIS Felt, 1908 robiniae (Haldeman, 1847) - Felixstowe, 12 November 2011, J.P. Bowdey, Host: Robinia pseudoacacia TM 24,33,36 [7] Unplaced genera of Cecidomyiinae DIODAULUS Rübsaamen, 1917 linariae (Winnertz, 1853) - Brockford, 1937, A. Mayfield, Host: Linaria vulgaris TM 16 [1] no recent records HARMANDIOLA Skuhravá, 1997 cavernosa (Rübsaamen, 1899) - Theberton Woods, 15 May 2014, J.P. Bowdey, Host: Populus tremula TM 46 [1] globuli (Rübsaamen, 1889) - Wherstead, 12 October 1974, A. Watchman, Host: Populus tremula TM 14,46 [2] tremulae (Winnertz, 1853) - Bentley Woods, no date known, C. Morley, Host: Populus tremula TM 05,13 [2] MASSALONGIA Kieffer, 1897 rubra (Kieffer, 1890) - Bradfield Woods, 1990-1991, B.R. Laurence, Host: Betula sp.TL 95 [1] MONARTHROPALPUS Rübsaamen, 1892 flavus (Schrank, 1776) - Ickworth, 19 September 2005, J.P. Bowdey, Host: Buxus sempervirens TL 86 [1] MYCOCECIS Edwards, 1922 ovalis Edwards, 1922 - Bradfield Woods, 14 August 1994, R. Hancy, Host: Hypoxylon sp. (fungi) TL 95 [1] PUTONIELLA Kieffer, 1896 pruni (Kaltenbach, 1872) - Monk Soham, no date known, C. Morley, Host: Prunus spinosa TM 26 [1] no recent records Supertribe Lasiopteridi BAYERIOLA Gagné, 1991 salicariae (Kieffer, 1888) - North Warren, Aldeburgh, 10 September 2019, J.P. Bowdey, Host: Lythrum salicaria TM 45 [1] DASINEURA Rondani, 1840 acrophila (Winnertz, 1853) - Bradfield Woods, 1990-1991, B.R. Laurence, Host: Fraxinus excelsior TL 95 TM 36,38,49,59 [7] angelicae Rübsaamen, 1916 - North Warren, Aldeburgh, 10 September 2019, J.P. Bowdey, Host: Angelica sylvestris TM 45 [1]

Trans. Suffolk Nat. Soc. 55 (2019)

Suffolk Natural History, Vol. 55 aparines (Kieffer, 1889) - Seafield Bay, Brantham, 14 June 2008, J.P. Bowdey, Host: Galium aparine TM 13,36 [2] crataegi (Winnertz, 1853) - Oulton Broad, 21 July 1990, P. Lee, Host: Crataegus monogyna TL 77,95 TM 03,05,07,13,14,15,23,25,36,37,46 [24] earlier records in TSNS without details fraxinea Kieffer, 1907 - Bradfield Woods, 1990-1991, B.R. Laurence, Host: Fraxinus excelsior TL 95 TM 03,25,37,48 [5] fraxini (Bremi, 1847) - Bradfield Woods, 1990-1991, B.R. Laurence, Host: Fraxinus excelsior TL 95 TM 04,15,25,36,37,46,59 [12] glechomae (Kieffer, 1889) - Shrubland Park, Barham, 6 October 1958, J.C.N. Willis, Host: Glechoma hederacea TL 94 TM 05,15,24,37 [6] gleditchiae (Osten Sacken, 1866) - Otley, 28 June 2010, J.P. Bowdey, Host: Gleditsia triacanthos TM 15,36,37,46 [6] hygrophila (Mik, 1883) - St Olaves, no date known, E.A. Ellis Host: Galium palustre TM 49 [1] no recent records kiefferiana (Rübsaamen, 1891) - Somerleyton, 15 August 1999, R. Hancy, Host: Chamerion angustifolium TM 24,37,49 [4] lathyri (Kieffer, 1909) - Kelsale, 13 June 2014, J.P. Bowdey, Host: Lathyrus pratensis TM 36 [1] lupulinae (Kieffer, 1891) - Kelsale, 4 August 2018, J.P. Bowdey, Host: Medicago lupulina TM 36 [1] marginemtorquens (Bremi, 1847) - Oulton Marsh, 23 September 1990. P. Lee Host: Salix viminalis TM 13,15,47.59 [4] odoratae Stelter,1982 - Holbrook, 3 June 2003, J.P. Bowdey, Host: Viola odorata TL 94 TM 13,15,23,33,34,36,37,44,57 [15] this species is yet to be added to the British checklist. It is probable that galls on V. odorata, recorded in Britain belong to D. odoratae but this requires confirmation (Chandler, 2019) plicatrix (Loew, 1850) - Somerleyton, 15 August 1999, R. Hancy, Host: Rubus fruticosus sp TG 40 TL 86 TM 03.36,45,46,47 49 [13] earlier records in TSNS without details populeti (Rübsaamen, 1889) - Dunwich Heath, 18 August 2017, BPGS meeting, Host: Populus tremula TM 46 [1] pteridis (Müller, 1871) - Herringfleet, July 1932, E.A. Ellis, Host: Pteridium aquilinum TG 40 TL 95 TM 46,47,49 [11] pustulans (Rübsaamen, 1889) - Southwold Covert, Wangford with Henham, 11 August 1991, P. Lee, Host: Filipendula ulmaria TL 95 TM 05,08,13,45,46,47,49 [10] pyri (Bouché, 1847) - Yoxford, 31 May 2015, J.P. Bowdey, Host: Pyrus domestica TM 36,38,46 [3] rosae (Bremi, 1847) - Trimley, July 1932, E.A. Ellis, Host: Rosa canina TL 94,98 TM 03,04,13,23,36,46 [14] strumosa (Bremi, 1847) - Bradfield Woods, 20 September 2014, BPGS meeting, Host: Lamiastrum galeobdolon TL 95 [1] thomasiana (Kieffer, 1888) - Yoxford, 18 November 2018, J.P. Bowdey, Host: Tilia sp. TM 36 [1]

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tiliae (Schrank, 1803) - Blythburgh, June 1934, E.A. Ellis, Host: Tilia sp. TM 46,47 [2] trifolii (F. Löw, 1874) - Flatford, 6 August 2008, BPGS meeting, Host: Trifolium repens TM 03,36,46 [4] tympani (Kieffer, 1909) - Cratfield, 13 September 2013, J. Ockenden, Host: Acer campestre TM 37 [1] ulmaria (Bremi, 1847) - Ickworth, 24 May 1988, D.K. Clements, Host: Filipendula ulmaria TL 77,84,86,95 TM 03,04,08,13,37,45,46,47,49,58 [25] ulmicola (Kieffer, 1909) - Somerleyton, 15 September 1999, R. Hancy, Host: Ulmus sp. TM 49 [1] urticae (Perris, 1840) - Barnby, 17 July 1991, P. Lee, Host: Urtica dioica TG 40 TL 86,95 TM 03,05,08,13,14,15,23,24,34,35,36,37,45,46,49,59 [37] earlier record in TSNS without details viciae (Kieffer, 1888) - Bradfield Woods, 14 August 1994, R. Hancy, Host: Vicia sp. TL 95 TM 13,36 [3] GEOCRYPTA Kieffer, 1913 galii (Loew, 1850) - Iken, 22 June 1980, E. Hyde, Host: Galium sp. TM 45,46 [3] GIRAUDIELLA Rübsaamen, 1915 inclusa (Frauenfeld, 1862) - Oulton Marshes, 31 December 1997, S. Youell, Host: Phragmites communis TM 13,46,59 [3] HARTIGIOLA Rübsaamen, 1912 annulipes (Hartig, 1839) - Shrubland Park, Barham, May-October 1995, C. Plant, Host: Fagus sylvatica TL 77,78,84 TM 14,15,23,36,46,59 [11] JAAPIELLA Rübsaamen, 1915 loticola (Rübsaamen, 1889) - Kelsale, 16 August 2016, J.P. Bowdey, Host: Lotus corniculatus TM 36 [1] parvula (Liebel, 1889) - Kelsale, 18 July 2018, J.P. Bowdey Host: Bryonia dioica TM 36 [1] veronicae (Vallot, 1827) - Iken, July 1932, E.A. Ellis, Host: Veronica chamaedrys TL 77,83,86,95,98 TM 03,04,05,07, 13,14,17,24,35,36,38,45,46,49,59 [52]. heraclei (Kaltenbach, 1862) - Flatford, 6 August 2008, BPGS meeting, Host: Heracleum sphondylium TM 03,15 [2] lamii Rübsaamen, 1916 - Needham Lake, 9 July 2011, BPGS meeting, Host: Lamium album TM 05 [1] lonicerae Rübsaamen, 1912 - Theberton Woods, 17 October 2015, J.P. Bowdey, Host: Lonicera periclymenum TM 46 [1] NEOMIKIELLA Hedicke, 1938 lychnidis (Vallot, 1827) - Sutton Common, 12 July 2019, L. Forsyth, Host: Silene latifolia TM 34 [1] RABDOPHAGA Westwood, 1847 heterobia (Loew, 1850) - Somerleyton, 15 August 1999, R. Hancy, Host: unknown TM 49 [1]

Trans. Suffolk Nat. Soc. 55 (2019)

Suffolk Natural History, Vol. 55 rosaria (Loew, 1850) - Barham, 15 April 1979, A. Watchman, Host: Salix sp. TL 77,84,87,94,95 TM 05,08,13,15,24,34,36,37,46,49 [21] earlier records in TSNS without details strobilina (Bremi, 1847) - Monk Soham, 23 May 1916, C. Morley, Host: Salix sp TL 94 TM 13,14,26 [4]

RONDANIOLA Rübsaamen & Hedicke, 1938 bursaria (Bremi, 1847) - Thornham Magna, 1920, C. Morley, Host: Glechoma hederacea TG 40 TL 86,94,95,98 TM 03,05,07,13,15,17,24,36,46,47,49 [26] WACHTLIELLA Rübsaamen, 1915 caricis (Loew, 1850) - Market Weston Fen, 7 July 2013, B. Eversham, Host: Carex disticha TL 95,97 [2] persicariae (Linnaeus, 1767) - Conard Mere, 12 August 1978, E. Milne-Redhead, Host: Persicaria sp TL 77,83 TM 03,08,13,35,36,37 [9] stachydis (Bremi, 1847) - Bradfield Woods, 20 September 2014, BPGS meeting, Host: Stachys sylvatica TL 95 TM 36,46,47 [4] Lasiopterini LASIOPTERA Meigen, 1818 carophila F. Löw, 1874 - Landguard Common, Felixstowe, 20 August 2015, J.P. Bowdey, Host: Daucus carota TM 23,36 [2] rubi (Schrank, 1803) - Bradfield Woods, 14 August 1994, R. Hancy, Host: Rubus sp. TG 40 TL 95,98 TM 04,13,46,47 [10] OZIRHINCUS Rondani, 1840 longicollis Rondani, 1840 - Knettishall Heath, 21 September 2014, BPGS meeting, Host: Tanacetum vulgare TL 98 TM 36 [3] Oligotrophini RHOPALOMYIA Rübsaamen, 1892 florum (Kieffer, 1890) - Flatford, 6 August 2008, BPGS meeting, Host: Artemisia vulgaris TM 07,13 [2] millefolii (Loew, 1850) - Knettishall Heath, 21 September 2014, BPGS meeting, Host: Achillea millefolium TL 86,98 TM 36 [3] tanaceticola (Karsch, 1879) - Middleton Moor, 27 July 2014, J.P. Bowdey, Host: Tanacetum vulgare TM 46 [2] Poomyini MAYETIOLA Kieffer, 1896 dactylidis Kieffer, 1896 - Kelsale, 5 November 2019, S. Carbonnelle, Host: Dactylis glomerata TM 36 [1] destructor (Say, 1817) - reference in TSNS without details Unplaced Lasiopteridi

CRANEIOBIA Kieffer, 1913 corni (Giraud, 1863) - Mendlesham, 1930-1939, A. Mayfield, Host: Cornus sanguinea TM 16 [1] no recent records Trans. Suffolk Nat. Soc. 55 (2019)

SUFFOLK DIPTERA CHECKLIST CYSTIPHORA Kieffer, 1892 sonchi (Vallot, 1827) - Sizewell, 12 September 1998, J.P. Bowdey, Host: Sonchus arvensis TL 94,96 TM 04,33,36,45,46,47 [12] DIDYMOMYIA Rübsaamen, 1912 tiliacea (Bremi, 1847) - Minsmere, 15 June 1990, E. Beaumont, Host: Tilia sp. TL 87 TM 04,14.15,36,46 [11] ITEOMYIA Kieffer, 1913 capreae (Winnertz, 1853) - Bradfield Woods, 14 August 1994, R. Hancy, Host: Salix sp. TL 94,95 TM 13,36,46,59 [9] major (Kieffer, 1898) - Bradfield Woods, 14 August 1994, R. Hancy, Host: Salix sp TL 86,94,95 TM 03,13,36,46,49,59 [12] MIKIOLA Kieffer, 1912 fagi (Hartig, 1839) - Stoke by Nayland, May 1935, F. Engleheart, Host: Fagus sylvatica TL 93 [1] no recent records

PHYSEMOCECIS Rübsaamen, 1914 ulmi (Kieffer, 1909) - Newbourne, 23 May 2015, R. Ruffell, Host: Ulmus sp. TM 24,36 [3] SEMUDOBIA Kieffer, 1913 betulae (Winnertz, 1853) - Belton Common, 15 August 1999, R. Hancy Host: Betula sp. TG 40 TM 05,13,45,46 [5] skuhravae Roskam, 1977 - Belton Common, 15 August 1999, R. Hancy Host: Betula sp. TG 40 TM 46 [2] tarda Roskam, 1977 - Needham Lake, 9 July 2011, BPGS meeting Host: Betula pendula TM 05,45 [2]

G. Calow

TAXOMYIA Rübsaamen, 1912 taxi (Inchbald, 1861) - Monks Eleigh, 18 July 1991, A. Watchman, Host: Taxus baccata TL 74,86,94 TM 03,14,25,36,37,38 [12]

Jaapiella veronicae galls on Germander Speedwell Trans. Suffolk Nat. Soc. 55 (2019)