The Diptera of Suffolk. by Suffolk Naturalists' Society

SUFFOLK DIPTERA CHECKLIST

THE DIPTERA OF SUFFOLK PETER VINCENT Introduction The records that form this checklist are from the Watsonian vice-counties of East (v.c. 25) and West (v.c. 26) Suffolk which are of greater extent than the modern administrative county boundaries. Diptera are not well recorded in Suffolk, with few records of even quite common species. Therefore, although records are in the main collected at monad (O.S.1km square) or greater accuracy, in order to assess the relative abundance and distribution of a diptera species the hectad (O.S.10km square) is used as the base recording unit. Suffolk comprises of 54 hectads or part hectads. In this checklist of Suffolk Diptera the arrangement of families follows as set out in Chandler (1998) and updates (Dipterists Forum, 2020). The classification of British diptera starts with the Lower Diptera (Nematocera) followed by the Brachycera. Within families, all taxa are listed alphabetically. Altogether there are at present 107 families of diptera recognised within the British fauna, and the British Isles list of diptera currently (April 2020) runs to some 7204 species (of which 41 are recorded from Ireland only) (Chandler, 2020). The layout of the checklist is that for each family the first division is by genus and then by species in alphabetical order. For most families, subfamilies and tribes are also recognised: and the genus can also be divided at subgenus level. For every individual species the species name (in bold type) is followed by the authority, then details of the first record of that species from Suffolk - place, date (to as much accuracy as possible) and recorder (where known). To give an indication of the distribution and the abundance of each species, this is followed by a list of all the hectads each species has occurred and the number of known records (in square brackets). Part Three: Psychodidae, Trichoceridae, Anisopodidae, Mycetobiidae, Scatopsidae, Ptychopteridae, Dixidae, Chaoboridae, Culicidae, Thaumaleidae, Simuliidae, Ceratopogonidae, Chironomidae. The checklist begins in Part One with the first infraorder of the Lower Diptera the Tipulomorpha followed in Part Two by the second infraorder of the Lower Diptera the Bibionomorpha. Part Three contains the remaining three infraorders of the Lower Diptera; Psychodomorpha, Ptychopteromorpha and Culicomorpha. The Psychodomorpha comprises of four superfamilies, Psychodoidea, Trichoceroidea, Anisopodoidea and Scatopsoidea. The Ptychopteromorpha comprises of one superfamily, Ptychopteroidea and the Culicomorpha comprises of two superfamilies, Culicoidea and Chironomoidea. In turn the Psychodoidea contains one family, Psychodidae. The Trichoceroidea contains one family, Trichoceridae, the Anisopodoidea contains two families, Anisopodidae and Mycetobiidae and the Scatopsoidea contains one family, Scatopsidae. The Ptychopteroidea contains one family Ptychopteridae. The Culicoidea contains three families, Dixidae, Chaoboridae and Culicidae, whereas the Chironomoidea contains four families, Thaumaleidae, Simuliidae, Ceratopogonidae and Chironomidae. Trans. Suffolk Nat. Soc. 56 (2020)

Suffolk Natural History, Vol. 56

Martin Cooper

Commonly known as moth flies or owl midges, Psychodidae, are a familiar and common family of flies. Despite being readily recognised by their small size, hairy wings and bead like antennae, they are difficult to identify to species level and are hence poorly recorded. Only 29 of 99 British species have been found in Suffolk. With slender bodies and long feeble legs Trichoceridae are similar in appearance to some of the smaller Tipulidae (craneflies). However, the presence of ocelli and differing wing venation distinguishes the Trichoceridae from these other families. They are one of very few insects that appear in adult form during winter months and often congregate into dancing swarms, even in Winter Gnat Trichocera regelationis quite cold weather; hence their popular name of winter gnats. Of the 10 British species 8 have been recorded from Suffolk. Anisopodidae and Mycetobiidae are two small families of flies. They are again slender, long legged flies. Until 1986 Mycetobiidae was included in Anisopodidae, but the two families can easily be separated by differing wing venation. Anisopodidae are commonly referred to as window gnats as they are familiar intruders to buildings; the adults often can be found sitting around windows and on walls where they are often mistaken for large mosquitoes (they cannot bite). There are 4 British species of Anisopodidae, 3 from Suffolk and 3 British species of Mycetobiidae of which there is only one Suffolk record. Scatopsidae are a family of very small to small black flies. They were once considered to be a subfamily of the Bibionidae, and in general appearance, short antennae and wing venation, that family does appear similar. However, there are more differences than similarities and are now not thought to be closely related at all. Scatopsidae are not easy to identify to species and although 26 of the 46 British species have been recorded from Suffolk there are comparatively few modern records from the county. Similar in appearance to some Tipulidae, Ptychopteridae are sometimes referred to by their common name of phantom craneflies and are one of the most striking and easily recognised of the Lower Diptera. They are medium sized to large flies with black lustrous bodies, often with yellow or brown markings on the abdomen and legs and with markings on the wings. Of the 7 British species 6 are found in Suffolk.

Trans. Suffolk Nat. Soc. 56 (2020)

Martin Cooper

Owl Midge Psychoda erminea

The three Culicoidea families of Dixidae, Chaoboridae and Culicidae are small slender flies, with narrow wings and delicate legs and antennae; all have aquatic larvae. Dixidae are commonly known as meniscus midges and Chaoboridae as phantom midges - so named because the larvae have a remarkably transparent body. Culicidae are the mosquitoes. Mosquitoes uniquely possess a long proboscis, which in the female, is adapted to penetrate the skin of an animal to permit blood-feeding, the blood used to provide protein for the development of their eggs. This blood feeding can be a nuisance to human activities as well as being the vector of several serious diseases around the world, of which malaria is the best known. There are 15 British Dixidae species, at present 11 of which have been found in Suffolk, 6 British Chaoboridae - 3 from Suffolk, and 32 British Culicidae - 24 from Suffolk. Thaumaleidae are small delicate yellowish-brown flies. They are very much associated with the upland areas of Northern and Western Britain, where the aquatic immature stages favour a very thin film of water flowing over stones or rocks; hence their common name of trickle midges. This type of habitat is rare in Suffolk and the only Suffolk record is of Thaumalea testacea Ruthe, 1831, where according to Claude Morley (1946-1948) ‘One female flew on to a motor car in Thelnetham Fen at 3 p.m. on the 25th June 1944’. There are 3 British species. Commonly known as black-flies, Simuliidae are small flies with broad wings, short stout bodies and short, thick legs. Both sexes often assemble in clouds around people’s heads, especially in sultry weather near water, the females’ bites being both irritating and capable of transmitting diseases. The immature stages are aquatic and require moving water. Many records in the A Black Fly Simuliidae checklist are of larvae and pupae. Of the 35 British species 18 have been recorded from Suffolk. The Ceratopogonidae are a large and difficult family with comparatively few Suffolk records. Though, anyone who has been to Scotland will be familiar with flies of this family, as the Highland midge Culicoides impunctatus Goetghebuer, 1920 belongs to the Ceratopogonidae or biting midges. They are very small flies, dark in colour, with broad wings. Females feed on the blood of vertebrates, including humans, to get protein for egg-laying and are responsible for the biological transmission of internationally important arboviruses of livestock. There have been 170 species recorded from Britain of which only 37 have been recorded from Suffolk. In many freshwater habitats, Chironomidae are among the most abundant flies, but again difficulties with identification mean they are poorly recorded, especially in Suffolk. In appearance Chironomidae are quite distinctive with a characteristically humped thorax that overhangs the head and in the main males have plumed antennae and wings that are shorter than the abdomen. Many chironomid species

Trans. Suffolk Nat. Soc. 56 (2020)

Paul Kitchener

SUFFOLK DIPTERA CHECKLIST

Suffolk Natural History, Vol. 56

show a wide range of tolerance to harsh environmental conditions and since chironomid larvae, are found in most of the types of freshwater ecosystems, they constitute good bioindicators to assess the water quality. Of the 610 British species 74 have been recorded from Suffolk. Chironomid Midge Records for this section of the checklist come mainly from Suffolk Biodiversity Information Centre (SBIS), and the National Biodiversity Network (NBN), with historical records from the Transactions of the Norfolk & Norwich Naturalists’ Society (Morley & Atmore, 1915) and the Transactions of the Suffolk Naturalists’ Society (Morley, 1946-1948). Other records were obtained from, Coe, Freeman & Mattingly (1950), Freeman & Lane (1985), Withers (1989), Laurence (1997), Snow (1998), and Crosskey (2005). References Chandler, P. J. (ed.) (1998). Checklists of Insects of the British Isles (New Series). Part 1; Diptera. Handbooks for the Identification of British Insects 12(1): 1-234. Chandler, P. J. (2020). Corrections and Changes to the Diptera Checklist (43). Dipterists Digest (Second Series) 27: 59-60. Coe, R. L., Freeman, P. & Mattingly, P. F. (1950). Diptera - 2. Nematocera: families Tipulidae to Chironomidae (Chironomidae). Handbooks for the Identification of British Insects 9 (2 iii): 121-246. Crosskey, R. W. (2005). A perspective on anthropophily in British blackflies (Diptera, Simuliidae), with keys for the identification of the culprit species. Dipterists Digest (Second Series) 12: 29-58. Dipterists Forum (2020). An Update of the 1998 Checklist of Diptera of the British Isles. Dipterists Forum [online] http://www. dipterists.org.uk/checklist/ [Accessed 9 October 2020] Freeman, P and Lane, R.P. (1985) Diptera: Bibionidae and Scatopsidae. Handbooks for the Identification of British Insects 9 (7); 26-45. Laurence, B. R. (1997). Flies from ancient coppiced woodland in Suffolk. Dipterist Digest (Second Series) 4: 78-91. Morley, C. & Atmore, E. A. (1915) The Diptera of Norfolk and Suffolk. Transactions of the Norfolk & Norwich Naturalists’ Society 10: 1-180. Morley, C. (1946-1948). The Diptera of Suffolk. Families Thaumaleidae, Anisopodidae, Psychodidae and Culicidae. Trans. Suffolk Nat. Soc. 6 (1): 3-8. Morley, C. (1946-1948). The Diptera of Suffolk. Families vii Chironomidae to xii Pulicidae. Trans. Suffolk Nat. Soc. 6 (2): 94-103. Snow, K. (1990). Mosquitoes. Naturalists’ Handbooks 14, Richmond Publishing Co. Ltd., Slough. Withers, P. (1989). Moth Flies Diptera: Psychodidae. Dipterist Digest 4: 1-81.

Trans. Suffolk Nat. Soc. 56 (2020)

Martin Cooper

SUFFOLK DIPTERA CHECKLIST

Suffolk Diptera Checklist (Part Three)

PSYCHODIDAE BOREOCLYTOCERUS Duckhouse, 1978 ocellaris (Meigen, 1804) - Lakenheath, 1983, R. Eley TL 68 TM 07,08 [6] PERICOMA Haliday in Walker, 1856 Subgenus PNEUMIA Enderlein, 1935 compta Eaton, 1893 - Bradfield Woods, October 1990, B.R. Laurence TL 95 [1] nubila (Meigen, 1818) - Redgrave & Lopham Fens, 10 June 2000, J. Cole TM 07,46 [2] pilularia Tonnoir, 1940 - Middleton, 17 October 2020, P.J. Vincent TM 46 [1] trivialis Eaton, 1893 - Monk Soham, 1945, C. Morley TL 86 TM 26,46 [5] Subgenus ULOMYIA Haliday in Walker, 1856 fuliginosa (Meigen, 1804) - Bury St Edmunds, 3 April 2006, B. Fountain TL 86 TM 46 [2] TONNOIRIELLA Vaillant, 1971 pulchra (Eaton, 1893) - Ickworh, 6 August 2003, D.J. Leeming TL 86 [6] Psychodini PSYCHODA Latreille, 1796 albipennis Zetterstedt, 1850 - Bradfield Woods, October 1990, B.R. Laurence TL 95 TM 07 [2] brevicornis Tonnoir, 1940 - no specific record details (Withers, 1989) cinerea Banks, 1894 - no specific record details (Withers, 1989) erminea Eaton, 1898 - Ipswich, 26 November 2013, M. Cooper TM 14 [1] grisescens Tonnoir, 1922 - no specific record details (Withers, 1989) minuta Banks, 1894 - Bradfield Woods, August 1991, B.R. Laurence TL 95 [1] phalaenoides (Linnaeus, 1758) - Wherstead, 21 April 1898, C. Morley TL 14 [1] setigera Tonnoir, 1922 - Bradfield Woods, August 1991, B.R. Laurence TL 95 [1] trinodulosa Tonnoir, 1922 - Minsmere, 2004, D. Gibbs TM 46 [1] TINEARIA Schellenberg, 1803 alternata (Say, 1824) - no specific record details (Withers, 1989)

Telmatoscopini PANIMERUS Eaton, 1913 albifacies (Tonnoir, 1919) - Redgrave & Lopham Fens, 1978-1985, P. Withers TM 07 [2] denticulatus Krek, 1972 - Suffolk - no specific record details (Withers, 1989) goetghebueri (Tonnoir, 1919) - Minsmere, 2004, D. Gibbs TM 46 [1] PARAMORMIA Enderlein, 1935 ustulata (Haliday in Walker, 1856 - Minsmere, 2004, D. Gibbs TM 46 [1] PERIPSYCHODA Enderlein, 1935 auriculata (Haliday in Curtis, 1839) - no specific record details (Withers, 1989) fusca (Macquart, 1826) - Ickworh, 6 August 2003, D.J. Leeming TL 86 [2]

Trans. Suffolk Nat. Soc. 56 (2020)

Suffolk Natural History, Vol. 56

PHILOSEPEDON Eaton, 1904 humeralis (Meigen, 1818) - Bradfield Woods, October 1990, B.R. Laurence TL 95 TM 46 [2] TELMATOSCOPUS Eaton, 1904 consors (Eaton, 1893) - Flixton, 14 August 1986, P. Withers TM 28,46 [2] parvulus Vaillant, 1960 - no specific record details (Withers, 1989) soleatus (Haliday in Walker, 1856 - Minsmere, 2004, D. Gibbs TM 46 [4] SYCORACINAE SYCORAX Haliday in Curtis, 1839 silacea Haliday in Curtis, 1839 - Redgrave & Lopham Fens, 1978-1985, P. Withers TM 07 [1] similis (Müller, 1927) - Redgrave & Lopham Fens, 23 June 2009, A. Gregory TM 07 [2]

TRICHOCERIDAE TRICHOCERA Meigen, 1803 Subgenus SALTRICHOCERA Krzemińska, 2002 annulata Meigen, 1818 - Beccles, January 1892, C. Morley TL 87,95 TM 46,49,57 [9] maculipennis Meigen, 1818 - Ipswich, 17 March 1895, C. Morley TM 14,26 [2] no recent records parva Meigen, 1804 - Barking Wood, October 1944, C. Morley TM 05,26,46 [3] regelationis (Linnaeus, 1758) - Ipswich, October 1895, C. Morley TL 67,95 TM 26,46 [8] rufescens Edwards, 1921 - Monk Soham, 1 May 1942, C. Morley TM 26 [2] no recent records saltator (Harris, 1776) - Bentley Woods, 1895, C. Morley TL 67,78 TM 13,46,58 [6] Subgenus TRICHOCERA hiemalis (De Geer, 1776) - Ipswich, December 1895, C. Morley TL 67,95 TM 14,46 [9] major Edwards, 1921 - Brandeston, 20 October 1943, C. Morley TM 26 [3] no recent records

ANISOPODIDAE SYLVICOLA Harris, 1780 cinctus (Fabricius, 1787) - Monk Soham, no date given, C. Morley TL 67,68 TM 14,33,46,47 [12] fenestralis (Scopoli, 1763) - Monk Soham, 29 June 1945, C. Morley TM 07,26 [2] punctatus (Fabricius, 1787) - Fritton, 1932, C. Morley TL 66,77,95 TM 07,15,34,46,49 [11]

MYCETOBIIDAE MYCETOBIA Meigen, 1818 pallipes Meigen, 1818 - Brandon, 9 June 1900, C. Morley & E.A. Elliot TL 78 [1] no recent records

Trans. Suffolk Nat. Soc. 56 (2020)

SUFFOLK DIPTERA CHECKLIST

SCATOPSIDAE ECTAETIINAE ECTAETIA Enderlein, 1912 clavipes (Loew, 1846) - Monk Soham, no date given, C. Morley TM 26 [1] no recent records PSECTROSCIARINAE ANAPAUSIS Enderlein, 1912 nigripes (Zetterstedt, 1860) - Dunwich Heath, 18 July 2003, M. Howe & L. Howe TM 46 [1] soluta (Loew, 1846) - Beccles, 5 August 1917, C. Morley TL 66,77,95 TM 16,49 [6] talpae (Verrall, 1912) - Barton Mills, June 1911, C.E. Collin TL 77 [1] no recent records SCATOPSINAE Colobostematini COLOBOSTEMA Enderlein, 1926 nigripenne (Meigen, 1830) - Monk Soham, 18 May 1940, C. Morley TM 26 [2] no recent records triste (Zetterstedt, 1850) - Bradfield Woods, July 1990, B.R. Laurence TL 77,95 [3] EFCOOKELLA Haenni, 1998 albitarsis (Zetterstedt, 1850) - Newmarket, no date given, C.E. Collin TL 77.95 TM 46 [3] FERNEIELLA Cook in Freeman, 1985 incompleta (Verrall, 1886) - Mildenhall, 1925, F.W. Edwards TL 77,97 [2] HOLOPLAGIA Enderlein, 1912 bullata (Edwards, 1925) - Thetford, 17 June 1880, G.H. Verrall TL 88 [1] first British record transversalis (Loew, 1846) - Dunwich Heath, 1 July 2013, I. Perry TM 46 [1] Rhegmoclematini PARASCATOPSE Cook, 1955 minutissima (Verrall, 1886) - Gisleham, 18 September 1929, C. Morley TM 44,58 [2] THRIPOMORPHA Enderlein, 1905 cooki (Hutson, 1970) - Middleton, 3 September 2019, P.J. Vincent TM 46 [1] coxendix (Verrall, 1912) - Barton Mills, 1912, G.H Verrall TL 77 TM 46 [3] halterata (Meigen, 1838) - Foxhall, 13 August 1904, C. Morley TM 24 [1] no recent records verralli (Edwards, 1934) - no specific record details (Freeman & Lane, 1985) Scatopsini APILOSCATOPSE Cook, 1974 bifilata (Haliday in Walker, 1856) - Monk Soham, 5 November 1944, C. Morley TM 26 [1] no recent records Trans. Suffolk Nat. Soc. 56 (2020)

Suffolk Natural History, Vol. 56 flavicollis (Meigen, 1818) - Tattingstone, 16 September 1903, C. Morley TM 13 [1] no recent records flavocincta (Duda, 1928) - no specific record details (Freeman & Lane, 1985) picea (Meigen, 1818) - Monk Soham, no date given, C. Morley TL 95 TM 26 [2] scutellata (Loew, 1846) - Blythburgh Woods, September 1913, C. Morley TM 47 [1] no recent records

REICHERTELLA Enderlein, 1912 geniculata (Zetterstedt, 1850) - Brandon, 20 June 1914, C. Morley TL 78 TM 16,26,46 [5] pulicaria (Loew, 1846) - Market Weston Fen, 25 May 2020, I. Perry TL 97 [1] SCATOPSE Geoffroy, 1762 notata (Linnaeus, 1758) - Ellough, 1 May 1939, C. Morley TM 48 [1] no recent records Swammerdamellini COBOLDIA Melander, 1916 fuscipes (Meigen, 1830) - Monk Soham, no date given, C. Morley TL 77 TM 26 [2] SWAMMERDAMELLA Enderlein, 1912 brevicornis (Meigen, 1830) - Southwold, September 1913, C. Morley TL 95 TM 46,57 [6]

PTYCHOPTERIDAE PTYCHOPTERA Meigen, 1803 albimana (Fabricius, 1787) - Barnby, 1898, C. Morley TL 86,97 TM 07,24,26,33,46,47,48,49,58,59 [30] contaminata (Linnaeus, 1758) - Orford, 1908, J.J.F.X. King TL 68,77,86 TM 04,07,14,27,28,33,34,36,44,45,46,47,48,49,59 [60] lacustris Meigen, 1830 - Ipswich, 1908, R. Bennett TM 07,14,24,44,46,47,49 [11] minuta Tonnoir, 1919 - Thorndon, 1 October 1944, C. Morley TL 86,97 TM 07,28,46,47,58,59 [50] paludosa Meigen, 1804 - Bentley Woods, 1922, C. Morley TM 07,13 [3] scutellaris Meigen, 1818 - Bentley Woods, 1 May 1897, C. Morley TL 84,86,97 TM 13,44,46,49 [12]

DIXIDAE DIXA Meigen, 1818 maculata Meigen, 1818 - Boyton, 1920, G.H. Verrall TL 75 TM 34 [3] nebulosa Meigen, 1830 - Fritton, March 1941, C. Morley TL 97 TM 34,46,49 [5] nubilipennis Curtis, 1832 - Nacton, 12 June 2016, A. Wood TM 14 [1] submaculata Edwards, 1920 - Flatford, 1971, G. Abbott & D.H.L. Disney TM 03 [1] DIXELLA Dyar & Shannon, 1924 aestivalis (Meigen, 1818) - Ipswich, 1908, R. Bennett TL 87,95 TM 03,46,57 [20] amphibia (De Geer, 1776) - Slaughden, Aldeburgh, 25 July 1899, C. Morley TL 86,97 TM03,07,45,46,47,57 [12] Trans. Suffolk Nat. Soc. 56 (2020)

SUFFOLK DIPTERA CHECKLIST

attica (Pandazis, 1933) - Flatford, 1 October 1971, G. Abbott & D.H.L. Disney TM 03,07.13,23,45,47,48,57 [30] autumnalis (Meigen, 1838) - Flatford, 1 October 1971, G. Abbott & D.H.L. Disney TL 77,86,97 TM 03,13,23,37,45,46,47,48,57 [51] graeca (Pandazis, 1937) - Walberswick, 11 February 1992, E. K. Goldie-Smith TM 48 [1] martinii (Peus, 1934) - Flatford, 1971, G. Abbott & D.H.L. Disney TL 97 TM 03,47 [3] serotina (Meigen, 1818) - Assington, 15 June 1900, C. Morley TL 77,88,97 TM 07,47 [12]

CHAOBORIDAE CHAOBORUS Lichtenstein, 1800 Subgenus CHAOBORUS sensu stricto crystallinus (De Geer, 1776) - Monk Soham, 24 May 1908, C. Morley TL 86 TM 07,26,37,46,59 [11] flavicans (Meigen, 1830) - Potters Bridge, Southwold, 27 July 1992, E.D. Taylor TL 86 TM 46,57 [6] Subgenus PEUSOMYIA Saether, 1970 pallidus (Fabricius, 1794) - Monk Soham, 2 May 1944, C. Morley TM 26,46 [2]

CULICIDAE ANOPHELES Meigen, 1818 Subgenus ANOPHELES sensu stricto atroparvus van Thiel, 1927* - Redgrave & Lopham Fens, 30 May 2010, A. Godfrey TM 07 [2] claviger (Meigen, 1804) - Foxhall, May 1900, C. Morley TL 86 TM 03,14,24,26,46 [10] messeae Falleroni, 1926* - Southwold, 1900, Critten TL 66,86 TM 05,14,16,17,23,24,27,34,35,44,46,47,57,59 [17] plumbeus Stephens, 1828 - Southwold, 14 June 1923, D.W. Collings TM 07,49,57 [4] CULICINAE AEDES Meigen, 1818 Subgenus AEDES sensu stricto cinereus Meigen, 1818 - Barton Mills, 1886, G.H. Verrall TL 77,78,94 [5] Subgenus AEDIMORPHUS Theobald, 1903 vexans (Meigen, 1830) - Tuddenham Fen, 24 August 1900, C. Morley TL 77,88 [2] no recent records COQUILLETTIDIA Dyar, 1905 richiardii (Ficalbi, 1889) - Southwold, 1924, D.W. Collings TM 37,46,47,57 [5]

*These two members of the A. maculipennis complex are morphologically identical except for the patterning on the decks of the eggs. Because of the difficulty in separating these species, most records, especially those pre-dating the recognition of the two species as distinct, are imprecise. (Snow, 1998)

Trans. Suffolk Nat. Soc. 56 (2020)

Suffolk Natural History, Vol. 56

CULEX Linnaeus, 1758 Subgenus CULEX sensu stricto pipiens Linnaeus, 1758 - Bentley Woods, 1 October 1898, C. Morley TL 66,77,86,94, TM 13,27 [12] pipiens form molestus Forskål, 1775 - Monk Soham, 10 October 1944, C. Morley TM 26 [3] no recent records Subgenus NEOCULEX Dyar, 1905 territans Walker, 1856 - Monk Soham, 31 March 1944, C. Morley TM 26 [1] no recent records

CULISETA Felt, 1904 Subgenus CULICELLA Felt, 1904 fumipennis (Stephens, 1825) - Mellis, 20 March 1946, Unknown TM 07 [1] no recent records litorea (Shute, 1928) - Southwold, 1 August 1900, D.W. Collings TM 47,57 [2] no recent records morsitans (Theobald, 1901) - Newmarket, 1908, G.H. Verrall TL 66 TM 46 [2] Subgenus CULISETA sensu stricto annulata (Schrank, 1776) - Bentley Woods, 1894, C. Morley TL 67,86 TM 07,13,17,26,37,46,49 [11] subochrea (Edwards in Wesenberg-Lund, 1921) - Monk Soham, 2 October 1944, C. Morley TM 26,58 [2] no recent records DAHLIANA Reinert, Harbach & Kitching, 2007 geniculata (Olivier, 1791) - Bentley Woods, 22 June 1945, C. Morley TL 95 TM 13 [3] OCHLEROTATUS Lynch Arribalzaga, 1891 Subgenus OCHLEROTATUS sensu stricto annulipes (Meigen, 1830) - Fritton, 20 March 1940, Dr Haynes TL 87 TM 07,46,47,49 [12] cantans (Meigen, 1818) - Barton Mills, 1916, G.H. Verrall - J.E. Collin TL 77,93,95 TM 46 [4] caspius (Pallas, 1771) - Southwold, 1924, D.W. Collings TL 78 TM 26,45,57 [6] communis (De Geer, 1776) - Monk Soham, 11 May 1945, C. Morley TM 26 [1] no recent records dorsalis (Meigen, 1830) - Aldeburgh, 1895, A. Piffard TL 77 TM 45,57 [3] no recent records flavescens (Müller, 1764) - Southwold, 20 September 1945, C. Morley TM 07,46,58 [3] punctor (Kirby in Richardson, 1837) - Blythburgh Woods, 14 September 1944, C. Morley TM 07,47 [2] sticticus (Meigen, 1838) - Monk Soham, 30 September 1944, C. Morley TM 26 [1] no recent records

Subgenus RUSTICOIDUS Shevchenko & Prudkina, 1973 rusticus (Rossi, 1790) - Monk Soham, no date given, C. Morley TM 26,35,37 [3]

Trans. Suffolk Nat. Soc. 56 (2020)

SUFFOLK DIPTERA CHECKLIST

THAUMALEIDAE THAUMALEA Ruthé, 1831 testacea Ruthé, 1831 - Thelnetham Fen, 25 June 1944, C. Morley TM 07 [1] no recent records

SIMULIIDAE SIMULIINAE Prosimuliini PROSIMULIUM Roubaud, 1906 tomosvaryi (Enderlein, 1921) - Washbrook, 28 March 2013, A. Chalkley TM 14 [1]

Simuliini SIMULIUM Latreille, 1802 Subgenus BOOPHTHORA Enderlein, 1921 erythrocephalum (De Geer, 1776) - Monk Soham, 24 May 1908, C. Morley TL 77 TM 26 [5] Subgenus EUSIMULIUM Roubaud, 1906 angustipes Edwards, 1915 - Boxted, 21 March 1993, A. Chalkley TL 93 TM 03,34 [3] aureum Fries, 1824 - Brightwell, 10 August 1992. B.A. Woodrow & A. Chalkley TM 24 [1] Subgenus HELLICHIELLA Rivosecchi & Cardinali, 1975 latipes (Meigen, 1804) - Brandon, 23 April 1904, C. Morley TL 78 TM 04,13,26,46 [7] Subgenus NEVERMANNIA Enderlein, 1921 angustitarse (Lundström, 1911) - Foxhall, 4 April 1896, C. Morley TL 93 TM 24,34 [4] costatum Friederichs, 1920 - Ipswich, 27 September 2000, P. Lee TL 93 TM 14 [6] cryophilum (Rubtsov, 1959) - Elmsett, 30 April 2013, A. Chalkley TM 04 [3] lundstromi (Enderlein, 1921) - Polstead, 1 June 2001, A. Chalkley TL 93 TM 03,04,36 [7] vernum complex Macquart, 1826 - sites in West Suffolk, 1 March 2001, A. Chalkley TL 64,65,74,84,85,93,94,95,97 [28] Subgenus SIMULIUM sensu stricto ODAGMIA Enderlein, 1921 argyreatum Meigen, 1838 - Elveden, 26 August 1992, A. Chalkley TL 67,78 TM 14,34 [4] intermedium Roubaud, 1906 - Shelley, 28 March 1993, A. Chalkley TM 03 [1] morsitans Edwards, 1915 - Fakenham, pre 1915, C.G. Nurse TL 97 [1] no recent records noelleri Friederichs, 1920 - Staverton Thicks, Wantisden, 18 August 1931, C. Morley TM 07,14,24,35,59 [7] ornatum complex Meigen, 1818 - Foxhall, 31 March 1902, C. Morley TL 85.93,94 TM 03,15,24,26 [13] posticatum Meigen, 1838 - Barnham, pre 1915, C.G. Nurse TL 77,85 TM 03 [4] reptans (Linnaeus, 1758) - Brandon, April 1904, C. Morley TL 79 TM 07,24 [5] variegatum Meigen, 1818 - Icklingham. 7 June 1947, C. Morley TL 77 [1] no recent records Trans. Suffolk Nat. Soc. 56 (2020)

Suffolk Natural History, Vol. 56

Subgenus WILHELMIA Enderlein, 1921 equinum (Linnaeus, 1758) - Barnham, Icklingham, West Stow, pre 1915, C.G. Nurse TL 77,87 [3] no recent records

CERATOPOGONIDAE CERATOPOGONINAE Ceratopogonini CERATOPOGON Meigen, 1803 lacteipennis Zetterstedt, 1838 - Minsmere, 6 July 2004, D. Agassiz TM 46 [1] SERROMYIA Meigen, 1818 femorata (Meigen, 1804) - Monk Soham, 26 July 1915, C. Morley TL 95 TM 26 [3] morio (Fabricius, 1775) - Bradfield Woods, 28 May 2009, I. Perry TL 95,97 [2] STILOBEZZIA Kieffer, 1911 Subgenus ACANTHOHELEA Kieffer, 1917 gracilis (Haliday, 1833) - Dunwich Heath, 25 June 2013, I. Perry TM 46 [1] Culicoidini CULICOIDES Latreille, 1809 Subgenus AVARITIA Fox, 1955 chiopterus (Meigen, 1830) - Monk Soham, 23 May 1916, C. Morley TM 26 [2] no recent records obsoletus (Meigen, 1818) - Monk Soham, 1 November 1922, C. Morley TL 95 TM 26,46 [4] Subgenus BELTRANMYIA Vargas, 1953 circumscriptus Kieffer, 1918 - Fritton Lake, 9 August 1934, C. Morley TM 49 [1] no recent records Subgenus CULICOIDES sensu stricto pulicaris (Linnaeus, 1758) - Oulton Broad, September 1913, C. Morley TL 95 TM 46,59 [3] Subgenus MONOCULICOIDES Khalaf, 1954 nubeculosus (Meigen, 1830) - Aldeburgh, 6 September 1907, G.H. Verrall TM 45 [1] no recent records Subgenus OECACTA Poey, 1851 minutissimus (Zetterstedt, 1855) - Monk Soham, no date given, C. Morley TM 26 [1] no recent records pictipennis (Staeger, 1839) - Wangford Wood, 12 May 1938, C. Morley TM 47 [1] no recent records vexans (Staeger, 1839) - Monk Soham, 14 August 1924, C. Morley TM 26 [1] no recent records Heteromyiini CLINOHELEA Kieffer, 1917 Subgenus CLINOHELEA sensu stricto unimaculata (Macquart, 1826) Trans. Suffolk Nat. Soc. 56 (2020)

SUFFOLK DIPTERA CHECKLIST

Palpomyiini BEZZIA Kieffer, 1899 Subgenus BEZZIA sensu stricto flavicornis (Staeger, 1839) - Dunwich Heath, 1 July 2013, I. Perry TM 46 [1] ornata (Meigen, 1830) - Easton Broad, 28 September 1923, C. Morley TM 46,57 [2] Subgenus HOMOBEZZIA Macfie, 1932 annulipes (Meigen, 1830) - Dunwich Heath, 1 July 2013, I. Perry TM 46 [1] Subgenus PYGOBEZZIA Remm, 1974 calceata (Haliday in Walker, 1856) - Aldeburgh, no date given, J.E. Collin TM 45,46 [3]

PALPOMYIA Meigen, 1818 armipes (Meigen, 1838) - Dunwich Heath, 8 Aug 2013, I. Perry TM 46 [1] distincta (Haliday, 1833) - Staverton Thicks, Wantisden, 23 June 1923, C. Morley TM 35 [1] no recent records flavipes (Meigen, 1804) - Tuddenham Heath, 1980-1990, Unknown TL 77 [1] grossipes Goetghebuer, 1920 - Dunwich Heath, 1 July 2013, I. Perry TM 46 [1] lineata (Meigen, 1804) - Market Weston Fen, 25 May 2010, I. Perry TL 97 TM 46 [2] luteifemorata Edwards, 1926 - Monk Soham, no date given, C. Morley TM 26 [1] no recent records serripes (Meigen, 1818) - Monk Soham, no date given, C. Morley TM 26 [1] no recent records spinipes (Meigen in Panzer, 1806) - Dunwich Heath, 25 June 2015, I. Perry TM46 [1] Sphaeromiini MALLOCHOHELEA Wirth, 1962 inermis (Kieffer, 1909) - Dunwich Heath, 25 June 2015, I. Perry TM46 [1] nitida (Macquart, 1826) - Dunwich Heath, 1 July 2013, I. Perry TM 46 [1] setigera (Loew, 1864) - Icklingham, 18 July 2000, Natural England TL 77 [1] SPHAEROMIAS Stephens in Curtis, 1829 pictus (Meigen, 1818) - Dunwich Heath, 8 August 2013, I. Perry TM 46 [1] DASYHELEINAE DASYHELEA Kieffer, 1911 Subgenus DASYHELEA sensu stricto flavifrons (Guérin-Ménéville, 1833) - Newmarket, July 1901, G.H. Verrall TL 66 TM 35 [2] no recent records Subgenus SEBESSIA Remm, 1979 acuminata Kieffer, 1919 - Monk Soham, 9 September 1920, C. Morley TM 26 [1] no recent records FORCIPOMYIINAE FORCIPOMYIA Meigen, 1818 Subgenus FORCIPOMYIA sensu stricto bipunctata (Linnaeus, 1767) - Monk Soham, 18 August 1917, C. Morley TM 26 [1] no recent records Trans. Suffolk Nat. Soc. 56 (2020)

100

Suffolk Natural History, Vol. 56

brevipennis (Macquart, 1826) - Ipswich, 1897, C. Morley TL 77 TM 26 [2] ciliata (Winnertz, 1852) - Monk Soham, no date given, C. Morley TM 26 [1] no recent records costata (Zetterstedt, 1838) - Newmarket, no date given, J.E. Collin TL 66 [1] no recent records myrmecophila (Egger, 1863) - Bentley Woods, March 1897, C. Morley TM 13,24 [2] no recent records nigra (Winnertz, 1852) - Newmarket, no date given, J.E. Collin TL 66 [1] no recent records radicicola Edwards, 1924 - Monk Soham, 24 May 1928, C. Morley TM 26 [2] no recent records

CHIRONOMIDAE CHIRONOMINAE Chironomini CHIRONOMUS Meigen, 1803 Subgenus CAMPTOCHIRONOMUS Kieffer, 1918 tentans Fabricius, 1805 - Newmarket, no date given, J.E. Collin TL 66 [1] no recent records Subgenus CHIRONOMUS sensu stricto aprilinus Meigen, 1830 - Southwold, April 1910, C. Morley TM 26,57 [2] no recent records plumosus (Linnaeus, 1758) - Monk Soham, 18 April 1914, C. Morley TM 17,26,46 [5] riparius Meigen, 1804 - Wherstead, no date given, C. Morley TM 14 [1] no recent records Subgenus LOBOCHIRONOMUS Ryser, Wülker & Scholl, 1985 dorsalis Meigen 1818 - Monk Soham, 11 May 1916, C. Morley TG 50 TM 07,26 [3] CLADOPELMA Kieffer, 1921 edwardsi (Kruseman, 1933) - Redgrave & Lopham Fen, 1978-85, P. Withers TM 07 [1] CRYPTOCHIRONOMUS Kieffer, 1918 redekei (Kruseman, 1933) - Redgrave & Lopham Fen, 31 May 2010, A. Godfrey TM 07 [1] supplicans (Meigen, 1830) - Aldeburgh, no date given, F. Jenkinson TM 45 [1] no recent records CRYPTOTENDIPES Beck & Beck, 1969 pseudotener (Goetghebuer, 1922) - Monk Soham, 22 May 1913, C. Morley TM 26 [1] no recent records DEMEIJEREA Kruseman, 1933 rufipes (Linnaeus, 1761) - Sudbury, no date given, B. Harwood TL 84 TM 14,26 [3] no recent records

Trans. Suffolk Nat. Soc. 56 (2020)

SUFFOLK DIPTERA CHECKLIST

101

DEMICRYPTOCHIRONOMUS Lenz, 1941 Subgenus DEMICRYPTOCHIRONOMUS sensu stricto vulneratus (Zetterstedt, 1838) - Monk Soham, 5 June 1919, C. Morley TM 26 [1] no recent records DICROTENDIPES Kieffer, 1913 nervosus (Staeger, 1839) - Monk Soham, May 1908, C. Morley TM 26 [1] no recent records EINFELDIA Kieffer, 1924 pagana (Meigen, 1838) - Monk Soham, 2 May 1944, C. Morley TM 26 [1] no recent records FLEURIA Kieffer, 1924 lacustris Kieffer, 1924 - Minsmere, 2004, D. Gibbs TM 46 [1] first British record GLYPTOTENDIPES Kieffer, 1913 Subgenus CAULOCHIRONOMUS Heyn, 1993 viridis (Macquart, 1834) - Monk Soham, 15 June 1931, C. Morley TM 26 [1] no recent records MICROTENDIPES Kieffer, 1915 chloris (Meigen, 1818) - Ipswich, April 1898, C. Morley TM 14 [1] no recent records pedellus (De Geer, 1776) - Southwold, 9 September 1921, C. Morley TL 78 TM 58 [2] no recent records PARATENDIPES Kieffer, 1911 albimanus (Meigen, 1818) - Monk Soham, June 1914, C. Morley TM 26,49 [2] no recent records PHAENOPSECTRA Kieffer, 1921 flavipes (Meigen, 1818) - Tuddenham Fen, September 1915, C. Morley TL 77,78 [2] no recent records POLYPEDILUM Kieffer, 1912 Subgenus POLYPEDILUM sensu stricto nubeculosum (Meigen, 1804) - Middleton, 12 April 2020, P.J. Vincent TM 46 [1] Subgenus TRIPODURA Townes, 1945 apfelbecki (Strobl, 1900) - Orford (Coe, Freeman & Mattingly, 1950) STENOCHIRONOMUS Kieffer, 1919 Subgenus PETALOPHOLEUS Borkent, 1984 fascipennis (Zetterstedt, 1838) - Newmarket, no date given, C.E. Collin TL 66 [1] no recent records SYNENDOTENDIPES Grodhaus, 1987 dispar (Meigen, 1830) - Ipswich, 1895, C. Morley TM 14,26 [2] no recent records

Trans. Suffolk Nat. Soc. 56 (2020)

102

Suffolk Natural History, Vol. 56

Tanytarsini MICROPSECTRA Kieffer, 1909 atrofasciata (Kieffer, 1911) - Redgrave & Lopham Fen, 1978-1985, P. Withers TM 07 [1] junci (Meigen, 1818) - Middleton, 31 March 2020, P.J. Vincent TM 46 [1] PARATANYTARSUS Thienemann & Bause, 1913 austriacus (Kieffer in Albrecht, 1924) - Redgrave & Lopham Fen, 1978-1985, P. Withers TM 07 [1] inopertus (Walker, 1856) - Mildenhall (Coe, Freeman & Mattingly, 1950) ORTHOCLADIINAE BRILLIA Kieffer, 1913 bifida (Kieffer, 1909) - Middleton, 23 October 2020, P.J. Vincent TM 46 [1] BRYOPHAENOCLADIUS Thienemann, 1934 tuberculatus (Edwards, 1929 - Bradfield Woods, May 1991, B.R. Laurence TL 95 [1] vernalis (Goetghebuer, 1921) - Bradfield Woods, October 1990, B.R. Laurence TL 95 [1] CAMPTOCLADIUS van der Wulp, 1874 stercorarius (De Geer, 1776) - Bentley Woods, July 1897, C. Morley TL 95 TM 13,14 [3] CHAETOCLADIUS Kieffer, 1911 perennis (Meigen, 1830) - Monk Soham, 18 April 1913, C. Morley TM 26 [1] no recent records piger (Goetghebuer, 1913) - Monk Soham, February 1938, C. Morley TM 26 [1] no recent records CORYNONEURA Winnertz, 1846 coronata Edwards, 1924 - Mildenhall, 25 September 1916, F.W. Edwards TL 77 [1] no recent records CRICOTOPUS van der Wulp, 1874 Subgenus CRICOTOPUS sensu stricto annulator Goetghebuer, 1927 - Newmarket, no date given, J.E. Collin TL 66 [1] no recent records bicinctus (Meigen, 1818) - Brandon, June 1914, C. Morley TL 78 [2] no recent records Subgenus ISOCLADIUS Kieffer, 1909 sylvestris (Fabricius, 1794) - Redgrave & Lopham Fen, 1978-1985, P. Withers TM 07 [1] tricinctus (Meigen, 1818) - Ipswich, 1895, C. Morley TM 14,48 [2] no recent records GYMNOMETRIOCNEMUS Edwards, 1932 brumalis (Edwards, 1929) - Bradfield Woods, October 1990, B.R. Laurence TL 95 TM 07,46 [3] subnudus (Edwards, 1929) - Bradfield Woods, 1977-1984, B.R. Laurence TL 95 [1]

Trans. Suffolk Nat. Soc. 56 (2020)

SUFFOLK DIPTERA CHECKLIST

103

HALOCLADIUS Hirvenoja, 1973 Subgenus HALOCLADIUS sensu stricto variabilis (Staeger, 1839) - Newmarket, no date given, J.E. Collin TL 66 [1] no recent records LIMNOPHYES Eaton, 1875 habilis (Walker, 1856) - Mildenhall, no date given, F.W. Edwards TL 77 [1] no recent records minimus (Meigen, 1818) - Newmarket, no date given, J.E. Collin TL 66 [1] no recent records

METRIOCNEMUS van der Wulp, 1874 albolineatus (Meigen, 1818) - Redgrave & Lopham Fen, 1978-1985, P. Withers TM 07 [1] eurynotus (Holmgren, 1883) - Bentley Woods, 28 March 1940, C. Morley TM 13 [1] no recent records fuscipes (Meigen, 1818) - Ipswich, 2 April 1895, C. Morley TM 14,26 [2] no recent records PARACLADIUS Hirvenoja, 1973 conversus (Walker, 1856) - Newmarket, no date given, G.H. Verrall TL 66 [1] no recent records

PARALIMNOPHYES Brundin, 1956 longiseta (Thienemann, 1919) - Redgrave & Lopham Fen, 1978-1985, P. Withers TM 07 [1] PSECTROCLADIUS Kieffer, 1906 Subgenus ALLOPSECTROCLADIUS Wülker, 1956 obvius (Walker, 1856) - Aldeburgh, no date given, F. Jenkinson TM 45 [1] no recent records PSEUDOSMITTIA Edwards, 1932 albipennis (Goetghebuer, 1921) - Bradfield Woods, May 1991, B.R. Laurence TL 95 [1]

SMITTIA Holmgren, 1869 aterrima (Meigen, 1818) - Monk Soham, 24 February 1939, C. Morley TL 95 TM 07,26 [3] contingens (Walker, 1856) - Redgrave & Lopham Fen, 1978-1985, P. Withers TM 07 [1] leucopogon (Meigen, 1804) - Redgrave & Lopham Fen, 1978-1985, P. Withers TM 07 [1] pratorum (Goetghebuer, 1927) - Redgrave & Lopham Fen, 1978-1985, P. Withers TM 07 [1] THIENEMANNIELLA Kieffer, 1911 majuscula (Edwards, 1924) - Mildenhall, 25 September 1916, F.W. Edwards TL 77 [1] no recent records

Trans. Suffolk Nat. Soc. 56 (2020)

104

Suffolk Natural History, Vol. 56

PRODIAMESINAE ODONTOMESA Pagast, 1947 fulva (Kieffer, 1919) - Bentley Woods, 3 May 1902, C. Morley TM 13 [1] no recent records Coelotanypodini CLINOTANYPUS Kieffer, 1913 Subgenus CLINOTANYPUS sensu stricto nervosus (Meigen, 1818) - Redgrave & Lopham Fen, 30 May 2010, A. Godfrey TM 07 [1]

Macropelopiini MACROPELOPIA Thienemann, 1916 nebulosa (Meigen, 1804) - Monk Soham, no date given, C. Morley TM 26 [1] no recent records notata (Meigen, 1818) - Sizewell, 9 September 1938, C. Morley TM 46 [1] no recent records PSECTROTANYPUS Kieffer, 1909 varius (Fabricius, 1787) - Monk Soham, 10 September 1945, C. Morley TM 07,26,46 [7] Natarsiini NATARSIA Fittkau, 1962 nugax (Walker, 1856) - Monk Soham, 21 May 1943, C. Morley TM 26,37 [2] no recent records punctata (Fabricius, 1805) - Monk Soham, 28 September 1940, C. Morley TM 26 [1] no recent records Pentaneurini ARCTOPELOPIA Fittkau, 1962 barbitarsis (Zetterstedt, 1850) - Mildenhall, F.W. Edwards (Transactions of the Entomological Society, 1929) CONCHAPELOPIA Fittkau, 1957 melanops (Meigen, 1818) - Newmarket, no date given, J.E. Collin TL 66 [1] no recent records MONOPELOPIA Fittkau, 1962 tenuicalcar (Kieffer, 1918) - Monk Soham, May 1908, C. Morley TM 26 [1] no recent records PARAMERINA Fittkau, 1962 cingulata (Walker, 1856) - Newmarket, no date given, J.E. Collin TL 66 [1] no recent records THIENEMANNIMYIA Fittkau, 1957 carnea (Fabricius, 1805) - Newmarket, no date given, J.E. Collin TL 66 [1] no recent records

Trans. Suffolk Nat. Soc. 56 (2020)

SUFFOLK DIPTERA CHECKLIST

105

TRISSOPELOPIA Kieffer, 1923 longimana (Staeger, 1839) - Redgrave & Lopham Fen, 1978-1985, P. Withers TM 07 [1] XENOPELOPIA Fittkau, 1962 falcigera (Kieffer, 1911) - Redgrave & Lopham Fen, 30 May 2010, A. Godfrey TM 07 [4] Procladiini PROCLADIUS Skuse, 1889 Subgenus HOLOTANYPUS Roback, 1982 choreus (Meigen, 1804) - Newmarket, no date given, J.E. Collin TL 66 [1] no recent records crassinervis (Zetterstedt, 1838) - Cavenham Heath, 31 July 1982, Unknown TL 77 TM 49 [2] culiciformis (Linnaeus, 1767) - Brandon, 25 August 1906, C. Morley TL 78 TM 57 [2] no recent records Subgenus PSILOTANYPUS Kieffer, 1906 lugens (Kieffer, 1915) - Timwoth, 1913, C.G. Nurse TL 86 [1] no recent records TELMATOGETONINAE THALASSOMYA Schiner, 1856 frauenfeldi Schiner, 1856 -Dunwich, 9 September 1921, C. Morley TG 50 TM 47 Peter Vincent Pennyfields Rectory Road Middleton Suffolk IP17 3NW

Trans. Suffolk Nat. Soc. 56 (2020)