Arabian Baboon (Papio hamadryas makkah Khalaf, 2015) A New Baboon Subsp. from Makkah, Saudi Arabia by Dr. Norman Ali Bassam Khalaf

The Arabian Baboon (Papio hamadryas makkah Khalaf, 2015) : A New Baboon Subspecies from Jabal Al-Noor in Makkah AlMukarramah, Kingdom of Saudi Arabia ‫ وُويع‬: )5102 ، ‫قرد الرباح العربي (بابيو هامادرياس مكت خلف‬ ‫ المملكت‬، ‫جديد مه قردة البابون مه جبل الىور في مكت ال ُمكرمت‬ ‫العربيت السعوديت‬ By: Prof. Dr. Sc. Norman Ali Bassam Ali Taher Khalaf-Sakerfalke von Jaffa

Arabian Baboon (Papio hamadryas makkah, Khalaf 2015) on Jabal Al-Noor in Makkah AlMukarramah, Kingdom of Saudi Arabia. Photo by: Prof. Dr. Sc. Norman Ali Bassam Khalaf-von Jaffa. 08.04.2012. http://www.flickr.com/photos/50022881@N00/11866019834/ .14..40.80 . ‫ نورمان علً بسام علً خلف الٌافاوي‬.‫ د‬.‫ أ‬: ‫ تصوٌر‬. ‫قرد الرباح العربً على جبل النور فً مكة المُكرمة‬

Abstract: A new subspecies of the Hamadryas Baboon from the genus Papio (Class Mammalia, Order Primates, Family Cercopithecidae) was found living on Jabal Al-Noor in Makkah Al-Mukarramah, Kingdom of Saudi Arabia. The new subspecies is distinguished from the five Papio hamadryas subspecies-groups: Papio hamadryas hamadryas (Sacred Gazelle : The Palestinian Biological Bulletin – Number 125 – May 2015

Baboon), Papio hamadryas anubis (Olive Baboon), Papio hamadryas cynocephalus (Yellow Baboon), Papio hamadryas papio (Red Baboon) and Papio hamadryas ursinus (Chacma Baboon), by its distinctive fur and face colouration and the moderate size. It is morphologically and geographically a distinct subspecies. The new subspecies was named Papio hamadryas makkah Khalaf, 2015.

Keywords: Mammalia, Primates, Cercopithecidae, Papio hamadryas makkah, Arabian Baboon, Makkah Baboon, Mecca Baboon, Jabal AlNoor, Mountain, Hijaz Mountains, Makkah, Makkah Al-Mukarramah, Mecca, Saudi Arabia, Kingdom of Saudi Arabia, Near East, Middle East.

‫ حٍث‬، ً‫ يتزا‬2.0 ‫ أػهى جبال يكت ٔػهى إرتفاع‬، ‫صؼدَا انٍٕو بحًد هللا تؼانى إنى لًت جبم انُٕر‬ ‫ ٔانذي أَشل ػُدِ انٕحً جبزٌم ػهٍّ انسالو أٔنى آٌاث انمزآٌ انكزٌى ػهى رسٕنُا‬، ‫ٌٕجد غار حِزاء‬ ‫ خهك‬.‫ إلزأ بإسى ربك انذي خهك‬.‫ بسى هللا انزحًٍ انزحٍى‬.‫انكزٌى يحًد صهى هللا ػهٍّ ٔسهى‬ ‫ صدق هللا انؼظٍى‬.‫ ػهى اإلَساٌ يا نى ٌؼهى‬.‫ انذي ػهى بانمهى‬.‫ الزأ ٔربك األكزو‬.‫اإلَساٌ يٍ ػهك‬ Family Khalaf (Prof. Dr. Norman, wife Ola and daughter Nora) climbed on 08.04.2012 Jabal Al-Noor, the highest mountain in Mecca (Elevation 642 meter) where they visited Ghar Hira' (Cave Hira'), the place where Prophet Mohammad Peace Be Upon Him, received the first revelation from ALLAH through the Messenger Angel Gabriel. https://www.flickr.com/photos/50022881@N00/17054529990/ Gazelle : The Palestinian Biological Bulletin – Number 125 – May 2015

I climbed Jabal Al-Noor (642 meter) in Makkah Al-Mukarramah, Kingdom of Saudi Arabia accompanied with my wife Ola Khalaf and daughter Nora Khalaf on 08.04.2012 during our Umrah visit in April 2012. Jabal Al-Noor (Mountain Al-Noor) belongs to the Hijaz Mountains. At the top, we visited Ghar Hira' (Cave Hira'), the place where Prophet Mohammad Peace Be Upon Him, received the first revelation from ALLAH through the Messenger Angel Gabriel. We saw many Arabian Baboons (Papio hamadryas makkah, Khalaf 2015) on Jabal Al-Noor in Makkah Al-Mukarramah. They were moving along the mountain stairs, and eating bananas given to them by the mountain climbers.

Arabian Baboon (Papio hamadryas makkah, Khalaf 2015) eating a banana on Jabal Al Noor in Makkah Al Mukarramah, Kingdom of Saudi Arabia. Photo by: Prof. Dr. Sc. Norman Ali Bassam Khalaf-von Jaffa. 08.04.2012. http://www.flickr.com/photos/50022881@N00/12064554125/

ً‫ نورمان (نعمان) علً بسام عل‬.‫ د‬.‫ أ‬: ‫ تصوٌر‬. ‫قرد الرباح العربً ٌأكل الموز على جبل النور فً مكة المُكرمة‬ .1....0.80 . ً‫طاهر خلف الٌافاوي العٌزري الفلسطٌنً األلمان‬

611 ‫ اآلٌة‬، ‫ سورة األعراف‬. ‫ٌِن‬ َ ‫َفلَمَّا َع َت ْوا َعنْ َما ُنهُوا َع ْن ُه قُ ْل َنا لَ ُه ْم ُكو ُنوا ق َِر َد ًة َخاسِ ئ‬

Hamadryas Baboon (Papio hamadryas Linnaeus, 1758) and the Arabian Baboon (Papio hamadryas makkah Khalaf, 2015) The Hamadryas Baboon (Papio hamadryas Linnaeus, 1758) is a species of baboon from the Old World monkey family. It is the northernmost of all the baboons, being native to the Horn of Africa and the southwestern tip of Gazelle : The Palestinian Biological Bulletin – Number 125 – May 2015

the Arabian Peninsula. These regions provide habitats with the advantage for this species of fewer natural predators than central or southern Africa where other baboons reside. The hamadryas baboon was a sacred animal to the ancient Egyptians and appears in various roles in ancient Egyptian religion, hence its alternative name of 'sacred baboon' (Wikipedia; Khalaf, August 2014).

.14..40.80 . ‫ نورمان علً بسام خلف‬.‫ د‬.‫ أ‬: ‫ تصوٌر‬. ‫قرد الرباح العربً على جبل النور فً مكة المُكرمة‬

‫ اآلٌة‬، ‫ سورة البقرة‬. ‫ٌِن‬ ِ ‫ٌِن اعْ َتد َْوا ِم ْن ُك ْم فًِ ال َّس ْب‬ َ ‫ت َفقُ ْل َنا لَ ُه ْم ُكو ُنوا ق َِر َد ًة َخاسِ ئ‬ َ ‫َولَ َق ْد َعلِ ْم ُت ُم الَّذ‬ . 16 Physical Description Apart from the striking size difference between the sexes (males are often twice as large as females), which is common to all baboons, this species also shows sexual dimorphism in coloration. The fur of males is silver-white in color and they have a pronounced cape (mane and mantle) which they develop around the age of ten, while the females are capeless and brown. Their faces range in color from red to tan to a dark brown (Wikipedia; Khalaf, August 2014). Males may have a body measurement of up to 80 cm (31 in) and weigh 20–30 kg (44–66 lb); females weigh 10–15 kg (22–33 lb) and have a body length of 40–45 cm Gazelle : The Palestinian Biological Bulletin – Number 125 – May 2015

(16–18 in). The tail adds a further 40–60 cm (16–24 in) to the length, and ends in a small tuft. Infants are dark in coloration and lighten after about one year. Hamadryas baboons reach sexual maturity at about 51 months for females and between 57 and 81 months for males (Wikipedia; Khalaf, August 2014).

َٰ َّ ‫َّللا ۚ َمنْ لَ َع َن ُه‬ ‫ب َعلَ ٌْ ِه َو َج َع َل ِم ْن ُه ُم ْالق َِر َد َة‬ َ ِ‫َّللا ُ َو َغض‬ ِ َّ ‫ك َم ُثو َب ًة ِع ْن َد‬ َ ِ‫قُ ْل َه ْل أ ُ َن ِّب ُئ ُك ْم ِب َشرٍّ ِمنْ َذل‬ َّ ‫ٌر َو َع َب َد‬ َ ‫الطا ُغ‬ ‫ اآلٌة‬، ‫ سورة المائدة‬. ‫ٌل‬ َ َ‫ك َشرٌّ َم َكا ًنا َوأ‬ َ ‫وت ۚ أُو َٰلَ ِئ‬ َ ‫از‬ ِ ِ‫ض ُّل َعنْ َس َوا ِء السَّب‬ ِ ‫َو ْال َخ َن‬ . 16

Ecology The baboon's range extends from the Red Sea in Eritrea to Ethiopia, Djibouti and Somalia. Baboons are also native to and live in southwestern and western Arabia, in both Yemen and Saudi Arabia. The Hamadryas baboon lives in semi-desert areas, savannas and rocky areas, requiring cliffs for sleeping and finding water. The hamadryas baboon is omnivorous and is adapted to its relatively dry habitat. During the wet seasons, the baboon feeds on a variety of foods, including blossoms, seeds, grasses, wild roots, and leaves from acacia trees. During the dry season, the baboons eat leaves of the Dobera glabra and sisal leaves. Hamadryas baboons also eat insects, reptiles and small mammals. One was even observed carrying a dead dik dik (Wikipedia; Khalaf, August 2014). The baboons’ drinking activities also depend on the season4 During the wet seasons, the baboons do not have to go far to find pools of water. During the dry Gazelle : The Palestinian Biological Bulletin – Number 125 – May 2015

seasons, they frequent up to three permanent waterholes. Baboons will take siestas at the waterholes during midafternoon. Hamadryas baboons will also dig drinking holes only a short distance from natural waterholes (Wikipedia; Khalaf, August 2014).

Social Life Group Organization The baboon has an unusual four-level social system called a multilevel society. Most social interaction occurs within small groups called one-male units or harems containing one male and up to ten females which the males lead and guard. A harem will typically include a younger "follower" male who may be related to the leader. Two or more harems unite repeatedly to form clans. Within clans, the dominant males of the units are probably close relatives of one another and have an age-related dominance hierarchy. Bands are the next level. Two to four clans form bands of up to 200 individuals which usually travel and sleep as a group. Both males and females rarely leave their bands. The dominant males Gazelle : The Palestinian Biological Bulletin â&#x20AC;&#x201C; Number 125 â&#x20AC;&#x201C; May 2015

will prevent infants and juveniles from interacting with infants and juveniles from other bands. Bands may fight with one another over food, etc., and the adult male leaders of the units are usually the combatants. Bands also contain solitary males that are not harem leaders or followers and move freely within the band. Several bands may come together to form a troop. Several bands in a troop also often share a cliff-face where they sleep (Wikipedia; Khalaf, August 2014).

Group Behaviour The hamadryas baboon is unusual among baboon and macaque species in that its society is strictly patriarchal. The males limit the movements of the females, herding them with visual threats and grabbing or biting any that wander too far away. Males will sometimes raid harems for females, resulting in aggressive fights. Many males succeed in taking a female from another's harem, called a "takeover". Visual threats are usually accompanied by these aggressive fights. This would include a quick flashing of the eyelids accompanied by a yawn to show off the teeth. As in many species, infant baboons are taken by the males as hostages during fights. However, males within the same clan tend to be related and respect the social bonds of their kin. In addition, females demonstrate definite preferences for certain males, and rival males heed these preferences. The less a female favors her harem males, the more likely she will be successfully taken by a rival. Young males, often "follower" males, may start their own harems by maneuvering immature females into following them. The male may Gazelle : The Palestinian Biological Bulletin â&#x20AC;&#x201C; Number 125 â&#x20AC;&#x201C; May 2015

also abduct a young female by force. Either way, the male will mate with the female when she matures. Aging males often lose their females to followers and soon lose weight and their hair color changes to brown like a female. While males in most other baboon species are transferred away from their male relatives and into different troops, male hamadryas baboons remain in their natal clans or bands and have associations with their male kin (Wikipedia). Hamadryas baboons have traditionally been thought of having a female transfer society with females being moved away from their relatives of the same sex. However, later studies show female baboons retain close associations with their female kin throughout their lives. Females can spend about as much time with other females as they do with the harem males, and some females will even interact with each other outside of their harems. In addition, it is not uncommon for females of the same natal group to end up in the same harem. Females can still associate and help their extended families despite their interactions being controlled by the harem males (Wikipedia). Females within a harem do not display any dominance relationships as seen in many other baboon and macaque species. The harem males suppress aggression between the females and prevent any dominance hierarchies from arising. Despite this, some social differences between the females occur. Some females are more socially active and have a stronger social bond with the harem male. These females, known as the "central females", stay in closer proximity to the harem male than the other females. Females that spend most their time farther from the harem male are called "peripheral females" (Wikipedia).

Reproduction and Parenting Like other baboons, the hamadryas baboon breeds aseasonally. The dominant male of a one-male unit does most of the mating, though other males may occasionally sneak in copulations, as well. Females do most of the parenting. They nurse and groom the infant and one female in a unit may groom an infant that is not hers. Like all baboons, hamadryas baboons are intrigued by their infants and give much attention to them. Dominant male baboons prevent other males from coming into close contact with their infants. They also protect the young from predators. The dominant male tolerates the young and will carry and play with them. When a new male takes over a female, she develops sexual swellings which may be an adaptation that functions to prevent the new male from killing the offspring of the previous male. When males reach puberty, they show interest in mothering young infants. They will kidnap the infants by luring them away from their harems and inviting them to ride on their backs. This is more often done by "follower" males. This kidnapping can lead to dehydration or starvation for the infant. The harem leader would retrieve the infants from their kidnappers, which is mostly an act to protect their offspring (Wikipedia).

Arabian Baboons (Papio hamadryas makkah, Khalaf 2015) on Jabal Al-Noor in Makkah AlMukarramah, Kingdom of Saudi Arabia. Photo by: Prof. Dr. Sc. Norman Ali Bassam Khalaf-von Jaffa. 08.04.2012. https://www.flickr.com/photos/50022881@N00/12064612585/

Gazelle : The Palestinian Biological Bulletin â&#x20AC;&#x201C; Number 125 â&#x20AC;&#x201C; May 2015

Human Interaction Cultural Depictions Hamadryas baboons often appear in ancient Egyptian art, as they were considered sacred to Thoth, a major and powerful deity with many roles that included being the scribe of the gods. Astennu, attendant to Thoth, is represented as a hamadryas in his roles as recorder of the result of the Weighing of the Heart and as one of the four hamadryas baboons guarding the lake of fire in Duat, the ancient Egyptian underworld. A pre-dynastic precursor to Astennu was Babi, or 'Bull of the Baboons', a bloodthirsty god said to eat the entrails of the unrighteous dead. Babi was also said to give the righteous dead continued virility, and to use his penis as the mast of a boat to convey them to the Egyptian paradise (Wikipedia). Sometimes Thoth himself appears in the form of a hamadryas (often shown carrying the moon on his head), as an alternative to his more common representation as an ibis-headed figure. Hapi, one of the Four Sons of Horus that guarded the organs of the deceased in ancient Egyptian religion, is also represented as hamadryas-headed: Hapi protected the lungs, hence the common Gazelle : The Palestinian Biological Bulletin â&#x20AC;&#x201C; Number 125 â&#x20AC;&#x201C; May 2015

sculpting of a stone or clay hamadryas head as the lid of the canopic jar that held the lungs and/or represented the protection of the lungs. Hamadryas baboons were revered because certain behaviors that they perform were seen as worshiping the sun (Wikipedia).

Status and Conservation Transformation of field and pastureland represents the main threat to the hamadryas baboon; it’s only natural predators are the Striped hyena, Spotted hyena and African leopard who are still living in its area of distribution (Wikipedia). In Arabia, the main enemy of the Arabian Baboon (Papio hamadryas makkah) is the rare Arabian Leopard (Panthera pardus nimr). The IUCN listed this species as "least concern" in 2008. There are no major rangewide threats at present, although locally it may be at risk through loss of habitat due to major agricultural expansion and irrigation projects. The species occurs in the proposed Yangudi Rassa National Park, the Harar Wildlife Sanctuary, and a number of wildlife reserves in the lower Awash valley and in northern Eritrea (Wikipedia). The Arabian Baboon (Papio hamadryas makkah) is also protected in the Yemeni Bur’ reserve and the Saudi Arabian reserves.

Conclusion: After studying and examining the Hamadryas Baboon Papio hamadryas from Jabal Al-Noor in Makkah Al-Mukarramah, and comparing with the five Papio hamadryas subspecies-groups: Papio hamadryas hamadryas (Sacred Baboon), Papio hamadryas Gazelle : The Palestinian Biological Bulletin – Number 125 – May 2015

anubis (Olive Baboon), Papio hamadryas cynocephalus (Yellow Baboon), Papio hamadryas papio (Red Baboon), Papio hamadryas ursinus (Chacma Baboon), and referring to many zoological references, and searching the Internet, I came finally to a conclusion that we are in front of a new Baboon subspecies from Jabal Al-Noor in Makkah Al-Mukarramah, Kingdom of Saudi Arabia. I gave it the scientific name Papio hamadryas makkah, new subspecies. The subspecies name “makkah” is for Makkah AlMukarramah, Saudi Arabia, where the new subspecies was discovered. Papio hamadryas makkah, new subspecies: Scientific trinomial name: Papio hamadryas makkah Khalaf, 2015 Authority: Prof. Dr. Sc. Norman Ali Bassam Khalaf-von Jaffa. Common Names: Arabian Baboon, Makkah Baboon, Mecca Baboon, Hijaz Mountains Baboon. Holotype: Phm-1, 40 cm, Dubai Zoo, Dubai, United Arab Emirates. Location: Jabal Al-Noor, Makkah Al-Mukarramah, Kingdom of Saudi Arabia. Date of Study: 8th April, 2012.

Gazelle : The Palestinian Biological Bulletin – Number 125 – May 2015

31 Photo from page 30 : Arabian Baboon (Papio hamadryas makkah, Khalaf 2015) on Jabal AlNoor in Makkah Al-Mukarramah, Kingdom of Saudi Arabia. Photo by: Prof. Dr. Sc. Norman Ali Bassam Khalaf-von Jaffa. https://www.flickr.com/photos/50022881@N00/12064635605/

https://www.flickr.com/photos/50022881@N00/12064924923/

http://www.flickr.com/photos/50022881@N00/11866268094/

Arabian Baboon (Papio hamadryas makkah, Khalaf 2015). Photo by the Author. http://www.flickr.com/photos/50022881@N00/11866213204/

Family Khalaf (Prof. Dr. Norman, wife Ola and daughter Nora) during their climbing on Jabal Al Noor in Makkah Al-Mukarramah, Kingdom of Saudi Arabia … The Holy City of Makkah (Mecca) is seen Behind them. 08.04.2012. https://www.flickr.com/photos/50022881@N00/16621913643/ ‫ َٕرياٌ (َؼًاٌ) ػهً بساو ػهً طاْز خهف انٍافأي انؼٍشري انفهسطًٍُ األنًاًَ يغ سٔجتّ انغانٍت‬.‫ د‬.‫أ‬ .1....0.80 . ‫ػال ٔإبُتّ انغانٍت َٕرة أثُاء صؼٕدْى ػهى جبم انُٕر ؛ ٔتتجهى يٍ ٔرائٓى يدٌُت يكت انًكزيت‬

References and Internet Websites: Berger, Lee R. and Brett Hilton-Barbe (2006). A Guide to Sterkfontein & the Cradle of Humankind. ISBN 978-1-77007-257-2. http://www.gettextbooks.com/isbn/9781770072572 BioLib. Biological Library. Taxon Profile Species Hamadryas Baboon Papio hamadryas (Linnaeus, 1758). http://www.biolib.cz/en/taxon/id32275/ Else, James G. and Phyllis C. Lee (Editors) (1986). Primate Evolution. New York: Cambridge University Press. 1986. xvii + 333 pp. http://onlinelibrary.wiley.com/doi/10.1002/ajpa.1330750119/abstract Khalaf, Norman Ali Bassam (891.)4 Tabie’t Al-Talawon fi Al-Haywanat (The Colouration of Animals). Al-Biology Bulletin. Number 1. January 1980, Safar 1401. Biological Society, Kuwait University, State of Kuwait. pp. 4-5. (In Arabic). Khalaf-von Jaffa, Norman Ali Bassam (1992). An Introduction to the Animal Life in Gazelle : The Palestinian Biological Bulletin – Number 125 – May 2015

35 Palestine. Gazelle. Gazelle: The Palestinian Biological Bulletin. Bonn-Bad Godesberg, Federal Republic of Germany. Number 30, Tenth Year, October 1992. pp. 1-7. (In Arabic). Khalaf-von Jaffa, Norman Ali Bassam (1994). An Introduction to the Animal Life in Palestine4 Shqae’q Al-Nouma’n (Anemone coronaria). A Quarterly Magazine Issued by the Program EAI (Education for Awareness and for Involvement). Environmental Education / Children for Nature Protection. In Cooperation with Dept. of General and Higher Education. P.L.O., Palestine. Number 4. Huzairan (June) 1994. pp. 16-21. (In Arabic). Khalaf-von Jaffa, Norman Ali (2004). Gazelle: Das Palästinensische Biologische Bulletin. Eine Wissenschaftliche Reise in Palästina, Arabien und Europa zwischen 1983 – 2004. / Gazelle: The Palestinian Biological Bulletin. A Scientific Journey in Palestine, Arabia and Europe between 1983 – 2004. ISBN 3-00-014121-9. Erste Auflage / First Edition, Juli 2004: 452 Seiten / Pages. Zweite erweiterte Auflage (Second Extended Edition), August 2004: 460 Seiten / Pages. Norman Ali Khalaf, Bonn-Bad Godesberg, Germany. http://drnorman-ali-khalaf-books.webs.com/ Khalaf, Norman Ali (2005, 2006, 2007). Chapter 3: Geography, Flora and Fauna. Pages 32-39. In: Palestine: A Guide. By Mariam Shahin, Photography by George Azar. CoAuthor: Norman Ali Khalaf. Northampton, Massachusetts: Interlink Publishing Group, 2005, 2006. xi + 471 pages. Appendices to page 500. http://ipsnewsite.mysite4now.com/journals.aspx?id=7323&jid=1&href=fulltext Khalaf-von Jaffa, Norman Ali (2006). Mammalia Arabica. Eine Zoologische Reise in Palästina, Arabien und Europa zwischen 1980-2006 / Mammalia Arabica. A Zoological Journey in Palestine, Arabia and Europe between 1980-2006. ISBN 3-00-017294-7. Erste Auflage, Juli 2006, 484 pp. Norman Ali Khalaf, Rilchingen-Hanweiler, Deutschland & Sharjah, United Arab Emirates. http://dr-norman-ali-khalafbooks.webs.com/mammaliaarabica.htm Khalaf-Sakerfalke von Jaffa, Norman Ali Bassam Ali Taher (2007). Haywanat Filistin (Fauna of Palestine). Wikipedia, Al-Mawsu'a Al-Hurra (The Free Encyclopedia). Gazelle: The Palestinian Biological Bulletin. Number 69, Twenty-fifth Year, September 2007 CE, Sha’ban 8.01 AH4 pp. 1-4. (in Arabic). http://ar.wikipedia.org/wiki/%D8%AD%D9%8A%D9%88%D8%A7%D9%86%D8%A7 %D8%AA_%D9%81%D9%84%D8%B3%D8%B7%D9%8A%D9%86 Khalaf-von Jaffa, Dr. Norman Ali Bassam (2009). Fauna Palaestina – Part One. A Zoological Journey in Palestine, Arabia and Europe between 1983 – 2006 / Fauna Palaestina – Teil Eins. Eine Zoologische Reise in Palästina, Arabien und Europa zwischen 1983 – 2006. ISBN 978-9948-03-865-8. Erste Auflage/First Edition, September 2009: 412 Seiten/Pages. Self Publisher: Dr. Norman Ali Bassam Khalaf-von Jaffa, Sharjah, United Arab Emirates & Rilchingen-Hanweiler, Bundesrepublik Deutschland. http://dr-norman-ali-khalaf-books.webs.com/faunapalaestinapart1.htm Khalaf-von Jaffa, Dr. Sc. Norman Ali Bassam Ali Taher (2010). Fauna Emiratus - Part One. Zoological Studies in the United Arab Emirates between 2004 - 2009. / Fauna Emiratus – Teil Eins. Zoologische Studien in die Vereinigten Arabischen Emirate zwischen 2004 - 2009. ISBN 978-9948-15-462-4. Erste Auflage/First Edition, November 2010: 350 Seiten / Pages. Self Publisher: Dr. Norman Ali Bassam Khalaf-von Jaffa, Dubai and Sharjah, United Arab Emirates & Rilchingen-Hanweiler, Bundesrepublik Deutschland. http://dr-norman-ali-khalaf-books.webs.com/faunaemiratuspart1.htm Khalaf-von Jaffa, Dr. Norman Ali Bassam (2012). Fauna Palaestina – Part Two. Gazelle : The Palestinian Biological Bulletin – Number 125 – May 2015

36 Zoological Studies in Palestine between 1983 – 2009 / Fauna Palaestina - Teil Zwei. Zoologische Studien in Palästina zwischen 1983 – 2009. ISBN 978-9948-16-667-2. 1. Auflage / First Edition : July 2012, Shaaban 1433 H. 208 Seiten / Pages (Arabic Part 120 Pages and the English Part 88 Pages). Publisher: Dar Al Jundi Publishing House, Jerusalem, Palestine. http://dr-norman-ali-khalafbooks.webs.com/faunapalaestinapart2.htm Khalaf-von Jaffa, Dr. Norman Ali Bassam (2013). Fauna Palaestina – Part Three. Zoological Studies in Palestine between 2005 – 2012 / Fauna Palaestina - Teil Drei. Zoologische Studien in Palästina zwischen 2005 – 2012. ISBN 978-9950-383-35-7. Erste Auflage / First Edition : July 2013, Shaaban 1434 H. 364 pages (English Part 350 Pages and the Arabic Part 14 Pages). Publisher: Dar Al Jundi Publishing House, Jerusalem, State of Palestine. http://dr-norman-ali-khalafbooks.webs.com/faunapalaestinapart3.htm Khalaf-von Jaffa, Prof. Dr. Norman Ali Bassam (2014). Fauna Palaestina – Part Four. Zoological Studies in Palestine between 1983 – 2014 / Fauna Palaestina - Teil Vier. Zoologische Studien in Palästina zwischen 1983 – 2014. ISBN 978-9950-383-77-7. Erste Auflage / First Edition : July 2014, Ramadan 1435 H. pp. 456 (English part 378 pages and Arabic part 78 pages). Publisher: Dar Al Jundi Publishing House, Al-Quds (Jerusalem), State of Palestine. http://fauna-palaestina-part-1.webs.com/faunapalaestina4.htm Khalaf-Sakerfalke von Jaffa, Prof. Dr. Sc. Norman Ali Bassam Ali Taher (2014). The Animals living inside and around the Sacred Mosque (Al-Masjid Al-Haram) in Makkah Al-Mukarramah, Kingdom of Saudi Arabia. Gazelle: The Palestinian Biological Bulletin. ISSN 0178 – 6288. Number 116, August 2014, Shawal 1435 AH. pp. 1-23. Dubai and Sharjah, United Arab Emirates. http://animals-of-makkah.webs.com/ Khalaf-Sakerfalke von Jaffa, Prof. Dr. Sc. Norman Ali Bassam Ali Taher (2015). The Arabian Baboon (Papio hamadryas makkah Khalaf, 2015) : A New Baboon Subspecies from Jabal Al-Noor in Makkah Al-Mukarramah, Kingdom of Saudi Arabia. Gazelle: The Palestinian Biological Bulletin. ISSN 0178 – 6288. Number 125, May 2015, Rajab 1436 AH. pp. 19-37. Dubai and Sharjah, United Arab Emirates. http://animals-ofmakkah.webs.com/arabian-baboon

Khalaf-von Jaffa, Dr. Norman Ali (2015). Taxon Profile Subspecies Papio hamadryas makkah Khalaf, 2015. BioLib. Biological Library. http://www.biolib.cz/en/adtaxon/id347918/ Khalaf-von Jaffa, Prof. Dr. Sc. Norman Ali Bassam (2015). Arabian Baboon (Papio hamadryas makkah Khalaf, 2015). EOL. Encyclopedia of Life. http://eol.org/collections/116554 Gazelle : The Palestinian Biological Bulletin – Number 125 – May 2015

37 Petraglia, Michael D. and Rose, Jeffrey I. (Eds.) (2009). The Evolution of Human Populations in Arabia: Paleoenvironments, Prehistory and Genetics. ISBN-13: 9789048127184. http://www.springer.com/gp/book/9789048127184 Wikipedia. Hamadryas Baboon. http://en.wikipedia.org/wiki/Hamadryas_baboon YouTube. Arabian Baboon Monkeys eating Banana on Jabal Al Noor in Mecca. 08.04.2012. By: Dr. Sc. Norman Ali Khalaf-von Jaffa. https://www.youtube.com/watch?v=pnLyCam4J50 YouTube. Arabian Baboon Monkeys along the roadside Makkah-Madinah Highway, Saudi Arabia. 10.04.2012. By: Dr. Sc. Norman Ali Khalaf-von Jaffa. https://www.youtube.com/watch?v=Frjwr8pQfQ8

Hamadryas Baboon (Papio hamadryas Linnaeus, 1758) at Dubai Zoo, Dubai, United Arab Emirates. Photo: Prof. Dr. Sc. Norman Ali Khalaf-von Jaffa. 11.03.2015. https://www.flickr.com/photos/50022881@N00/16596211537/ Gazelle : The Palestinian Biological Bulletin – Number 125 – May 2015