1
A Rare Stamp from Palestine: The Rockhopper Penguin (Eudyptes chrysocome) & Gentoo Penguin (Pygoscelis papua) Stamp By: Sharif Prof. Dr. Sc. Norman Ali Bassam Ali Taher Mohammad Ahmad Ahmad Mostafa Abdallah Mohammad Khalaf-Prinz Sakerfalke von Jaffa
In 2007, The Palestinian National Authority, State of Palestine, issued a rare post stamp showing a Rockhopper Penguin (Eudyptes chrysocome) in the main 1000 Mils or Fils stamp, and a Gentoo Penguin (Pygoscelis papua) with chick. The Rockhopper Penguins are divided into 3 subspecies: The Southern Rockhopper: Eudyptes chrysocome chrysocome, Eastern Rockhopper: Eudyptes chrysocome filholi and the Northern Rockhopper: Eudyptes chrysocome moseleyi. Gazelle : The Palestinian Biological Bulletin – Number 181 – January 2020
2
Breeding Range: Southern Rockhopper: Falkland Islands, Argentina and Chile. Eastern Rockhopper: Marion, Prince Edward, Crozet, Kerguelen, Heard, MacDonald, Macquarie, Campbell, Antipodes and Auckland Islands Northern Rockhopper: Tristan de Cunha, Gough, St Paul and Amsterdam Islands (Bingham). Length: 52 cm. Weight: 3 Kg. World Population: Southern Rockhopper: 650,000 breeding pairs Eastern Rockhopper: 800,000 breeding pairs Northern Rockhopper: 350,000 breeding pairs (Bingham). World-wide there are 3 subspecies of Rockhopper penguin; Southern, Eastern and Northern. Rockhopper penguins are amongst the smallest of the world's penguins, having an average length of around 52cm, and an average weight of about 3kg. A yellow stripe above each eye projects into a yellow crest, and these are joined behind the head by a black occipital crest. The eyes are red, the short bulbous bill is reddish brown, and the feet and legs are pink. The females are slightly smaller than the males, but have similar plumage (Bingham). Rockhopper penguin breeding colonies may be very large, with up to a hundred thousand nests at a single breeding site. Nesting densities range from 1.5 to 3 nests per sq.m., and colonies are often shared with nesting albatross or cormorants. Rockhopper penguins not only return to the same breeding site each year, but they also utilise the same nest, which they refurbish with stones, sticks, vegetation or any other suitable material (Bingham). The preferred nesting sites are steep rocky gullies, above approaches into deep water. Such sites may be vegetated by grasses or dwarf shrubs, but long-established colonies will generally have destroyed most of the natural vegetation surrounding the colony, and worn a pathway from the sea up the rock-face. Rockhopper penguins regularly bathe and drink fresh water, and most breeding sites are close to natural springs or freshwater puddles (Bingham). The breeding cycle begins in early October, with males arriving at the breeding site a few days earlier than the females. Copulation begins as soon as the females arrive, and egg-laying takes place in early November. Two eggs are laid 4 - 5 days apart, with the first egg hatching after the second. The first egg, at around 80g, is considerably smaller than the second egg of around 110g. This strategy aims to rear just one healthy chick under a wide range of circumstances. The second egg is generally brooded at the rear,
Gazelle : The Palestinian Biological Bulletin – Number 181 – January 2020
3
where the temperature is more stable, and where it is less prone to being lost or stolen (Bingham). In Eastern and Northern Rockhopper penguins it is almost unheard of for both chicks to be reared, but the Southern Rockhopper penguin is capable of rearing both chicks to fledging when conditions are favourable. Even so, Southern Rockhopper penguins show little annual fluctuation in population size and chick rearing success, and annual breeding success never exceeds 1 chick per breeding pair for the colony as a whole (Bingham). Incubation of the eggs takes around 33 days, and is divided into three roughly equal shifts. During the first shift both parents are in attendance. The male then goes to sea to feed while the female takes the second shift, and he returns to relieve the female for the third shift. The male remains on the nest until the eggs hatch, and continues to brood for the first 25 days, while the female brings food for the chicks (Bingham). Such a system of extended shift duration requires lengthy fasts for both parents, but allows them to forage further afield than would be the case if they had a daily changeover. The newly hatched chicks may have to wait for up to a week before the female returns with their first feed. During this period chicks are able to survive on existing yolk reserves, after which they begin receiving regular feeds of around 150g in weight (Bingham). By the end of the 25 days of brooding, chicks have developed their mesoptile plumage, and are receiving regular feeds averaging around 600g. By this stage they are able to leave the nest and creche with other chicks, allowing both adults to forage to meet the chicks' increasing demands for food. Rockhopper penguin creches are not generally as large as for species such as Gentoo penguins, possibly due to the more rugged terrain, and chicks creche into numerous small groups scattered throughout the colony (Bingham). Chicks completely lack the yellow markings of the adult birds, and even the bills are black. As chicks moult into adult plumage, the colony is joined by pre-breeding birds arriving to moult. These birds are distinguished from newly fledged chicks by a faint yellow stripe above the eye, and a reddish brown bill. The crest is not developed until birds mature. Rockhoppers do not breed until at least 4 years of age, but have been shown to live for up to 25 years in captivity (Bingham). Despite being amongst the smallest of penguins, Rockhopper penguins are perhaps the most aggressive. They show little fear of people, or of birds and animals larger than themselves. Anything that comes within range of an incubating bird will be pecked, including another Rockhopper, or the long wings of neighbouring albatross.
Gazelle : The Palestinian Biological Bulletin – Number 181 – January 2020
4
Nevertheless, Rockhopper penguins can be very gentle with their partners, and allopreening (mutual preening) is common (Bingham). Chicks fledge at around 10 weeks of age, and adults then spend 20 - 25 days at sea building up subcuticular body fat reserves in preparation for their annual moult. The moult lasts for around 25 days, and the birds then abandon the breeding site and spend the winter feeding at sea, prior to returning the following spring (Bingham). Rockhopper penguins are opportunistic feeders, and around the Falklands are known to take varying proportions of crustaceans (Euphausia lucens, E. vallentini, Thysanoessa gregaria and Themisto sp.), squid (Gonatus antarcticus, Loligo gahi, Onychoteuthis sp., and Teuthowenia sp.) and various small fish. Foraging dives rarely exceed 100m in depth, but feeding in groups is common (Bingham).
Rockhopper Penguin (Eudyptes chrysocome). Photo by Robert Orr. https://lizbrownleepoet.com/2016/04/21/r-is-for-rockhopper-penguin/
The Falkland Islands used to be the world's largest Rockhopper penguin breeding site, with an estimated 2,500,000 breeding pairs in 1984. This has since declined to around 300,000 breeding pairs as a result of commercial fishing around the Falklands (See Bingham 2002, Revista Chilena de Historia Natural 2002: The decline of Falkland Islands penguins in the presence of a commercial fishing industry). The same subGazelle : The Palestinian Biological Bulletin – Number 181 – January 2020
5
species of Rockhopper penguins in nearby South America have not declined, and there is some evidence of possible emigration from the Falklands to Staten Island in Argentina (Bingham). Eastern Rockhopper penguins have also undergone major declines over recent years, and although commercial fisheries operate in the areas of decline, no definite link has yet been established. Northern Rockhopper penguins have also declined, and in this instance commercial fishing with gill-nets, and egging, have been identified as the major causes (Bingham). The large scale decline in Rockhopper penguin populations has been of such magnitude as to justify treating the species as globally threatened according to the new IUCN criteria. In September 2001 the IPCWG recommended the exclusion of large scale commercial fishing from within 30 miles of penguin breeding sites in the Falkland Islands. So far the Falkland Islands Government has refused to adopt this measure, and penguins continue to decline as a result (Bingham). The Gentoo Penguin (Pygoscelis papua) is a penguin species in the genus Pygoscelis, most closely related to the Adélie penguin (P. adeliae) and the chinstrap penguin (P. antarcticus). The earliest scientific description was made in 1781 by Johann Reinhold Forster with a reference point of the Falkland Islands. They call in a variety of ways, but the most frequently heard is a loud trumpeting which the bird emits with its head thrown back (Wikipedia). The application of gentoo to the penguin is unclear. Gentoo was an Anglo-Indian term to distinguish Hindus from Muslims. The English term may have originated from the Portuguese gentil ("pagan, gentile"). Some speculate that the white patch on the bird's head was thought to resemble a turban (Wikipedia). It may also be a variation of another name for this bird, "Johnny penguin", Johnny being Juanito in Spanish and sounds vaguely like gentoo. The Johnny rook, a predator, is likely named after the Johnny penguin (Wikipedia). The specific name papua is a misnomer; in the original description, Johann Reinhold Forster, a naturalist who had circumnavigated the world with Captain James Cook, mistakenly assumed that the species occurred in Papua (New Guinea), the closest gentoos actually being over 6000 km to the south (on Macquarie Island). There are no penguins in New Guinea, however. Others trace the error to a "possibly fraudulent claim" in 1776 by French naturalist Pierre Sonnerat, who also alleged a Papuan location for the king penguin despite never having been to the island himself (Wikipedia). The gentoo penguin is one of three species in the genus Pygoscelis. Mitochondrial and nuclear DNA evidence suggests the genus split from other penguins around 38 million years ago, about 2 million years after the ancestors of the genus Aptenodytes. In turn, the Adélie penguins split off from the other members of the genus around 19 million years Gazelle : The Palestinian Biological Bulletin – Number 181 – January 2020
6
ago, and the chinstrap and gentoo finally diverged around 14 million years ago (Wikipedia). Two subspecies of this penguin are recognised: Pygoscelis papua papua and the smaller Pygoscelis papua ellsworthi (Wikipedia). The gentoo penguin is easily recognized by the wide white stripe extending like a bonnet across the top of its head and its bright orange-red bill. It has pale whitish-pink webbed feet and a fairly long tail – the most prominent tail of all penguin species. Chicks have grey backs with white fronts. As the gentoo penguin waddles along on land, its tail sticks out behind, sweeping from side to side, hence the scientific name Pygoscelis, which means "rump-tailed" (Wikipedia).
A Gentoo Penguin (Pygoscelis papua) parent feeding chick. Photo taken by Ian Wood on 24 January 2009. https://www.hbw.com/ibc/photo/gentoo-penguin-pygoscelis-papua/parentfeeding-chick
Gentoos reach a height of 51 to 90 cm (20 to 35 in), making them the third-largest species of penguin after the emperor penguin and the king penguin. Males have a maximum weight of about 8.5 kg (19 lb) just before molting, and a minimum weight of about 4.9 kg (11 lb) just before mating. For females, the maximum weight is 8.2 kg (18 lb) just before molting, but their weight drops to as little as 4.5 kg (9.9 lb) when Gazelle : The Palestinian Biological Bulletin – Number 181 – January 2020
7
guarding the chicks in the nest. Birds from the north are on average 700 g (1.5 lb) heavier and 10 cm (3.9 in) taller than the southern birds. Southern gentoo penguins reach 75–80 cm (30–31 in) in length. They are the fastest underwater swimmers of all penguins, reaching speeds of up to 36 km/h (22 mph). Gentoos are well adapted to extremely cold and harsh climates (Wikipedia). The breeding colonies of gentoo penguins are located on ice-free surfaces. Colonies can be directly on the shoreline or can be located considerably inland. They prefer shallow coastal areas and often nest between tufts of grass. In South Georgia, for example, breeding colonies are 2 km inland. Whereas in colonies farther inland, where the penguins nest in grassy areas, they shift location slightly every year because the grass will become trampled over time (Wikipedia). Gentoos breed on many sub-Antarctic islands. The main colonies are on the Falkland Islands, South Georgia and the South Sandwich Islands, and Kerguelen Islands; smaller colonies are found on Macquarie Island, Heard Islands, South Shetland Islands, and the Antarctic Peninsula. The total breeding population is estimated to be over 600,000 birds. Gentoos breed monogamously, and infidelity is typically punished with banishment from the colony. Nests are usually made from a roughly circular pile of stones and can be quite large, 20 cm (7.9 in) high and 25 cm (9.8 in) in diameter. The stones are jealously guarded and their ownership can be the subject of noisy disputes and physical attacks between individuals. They are also prized by the females, even to the point that a male penguin can obtain the favors of a female by offering her a choice stone (Wikipedia). Two eggs are laid, both weighing around 130 g (4.6 oz). The parents share incubation, changing duty daily. The eggs hatch after 34 to 36 days. The chicks remain in the nests for around 30 days before joining other chicks in the colony and forming crèches. The chicks molt into subadult plumage and go out to sea at around 80 to 100 days (Wikipedia). Gentoos live mainly on crustaceans, such as krill, with fish making up only about 15% of the diet. However, they are opportunistic feeders, and around the Falklands are known to take roughly equal proportions of fish (Patagonotothen sp., Thysanopsetta naresi, Micromesistius australis), squat lobsters (Munida gregaria), and squid (Loligo gahi, Gonatus antarcticus, Moroteuthis ingens) (Wikipedia). The gentoos' diet is high in salt as they eat organisms with relatively the same salinity as sea water, and this can lead to complications associated with high sodium concentrations in the body, especially for gentoo chicks. To counteract this, gentoos as well as many other marine bird species have a highly developed salt gland located above their eyes that takes the high concentration of sodium within the body and produces a highly saline-concentrated solution that drips out of the body from the tip of the beak (Wikipedia). Gentoo penguins do not store as much fat as the Adélie penguin, their closest relative: gentoos require less energy investment when hunting because the net gain of energy Gazelle : The Palestinian Biological Bulletin – Number 181 – January 2020
8
after hunting is greater in gentoos than Adélies. As embryos, gentoos require a lot of energy in order to develop. Oxygen consumption is high for a developing gentoo embryo. As the embryo grows and requires more oxygen, the amount of consumption increases exponentially until the gentoo chick hatches. By then, the chick is consuming around 1800 mL O2 per day (Wikipedia). In the sea, leopard seals, sea lions, and killer whales are all predators of the gentoo. On land, no predators of full-grown healthy gentoo penguins exist. Skuas and giant petrels regularly kill many chicks and steal eggs; Petrels will kill injured and sick adult gentoos. Various other seabirds, such as the kelp gull and snowy sheathbill, will also snatch chicks and eggs. Skuas on King George Island have been observed attacking and injuring adult gentoo penguins in apparent territorial disputes (Wikipedia). As of 2019, the IUCN Red List lists the gentoo as least concern, although rapid declines in some key areas are believed to be driving a moderate overall decline in the species population. Examples include Bird Island, South Georgia, where the population has fallen by two-thirds over 25 years (Wikipedia).
References and Internet Websites Bingham, Mike. Rockhopper Penguin. Organisation for the Conservation of Penguins, Argentina and Chile. http://www.penguins.cl/rockhopper-penguins.htm Bingham, Mike (2002). The decline of Falkland Islands penguins in the presence of a commercial fishing industry. Rev. chil. hist. nat. Revista chilena de historia natural. 2002, vol.75, n.4, pp.805-818. ISSN 0716-078X. http://dx.doi.org/10.4067/S0716078X2002000400014 & https://scielo.conicyt.cl/scielo.php?script=sci_abstract&pid=S0716078X2002000400014&lng=es&nrm=iso&tlng=en Brownlee, Liz (April 2016). R is for Rockhopper Penguin. https://lizbrownleepoet.com/2016/04/21/r-is-for-rockhopper-penguin/ Dardona, Ayman Wadi’ Youssef (M.Sc. Limnology) and Prof. Dr. Sc. Norman Ali Bassam Khalaf-von Jaffa (Dr. Sc. Zoology) (2015). Studying Aquatic Birds in the area between the Gaza Fishermen Port and Wadi Gaza Estuary, Gaza Strip, Palestine. Gazelle: The Palestinian Biological Bulletin. ISSN 0178 – 6288. Number 124, April 2015. pp. 22-39. Dubai and Sharjah, United Arab Emirates. http://birds-ofpalestine.webs.com/gaza-aquatic-birds Khalaf, Nora Norman Ali Bassam (Fotografin, 11 Jahre) (28 Juni 2010). Foto: Junger Strauß, Vogel und Tier Markt, Sharjah, Vereinigte Arabische Emirate. Fotocommunity.www.fotocommunity.de/pc/pc/display/21915604 Khalaf, Norman Ali Bassam (1980). Tabie’t Al-Talawon fi Al-Haywanat (The Colouration of Animals). Al-Biology Bulletin. Number 1. January 1980, Safar 1401. Biological Society, Kuwait University, State of Kuwait. pp. 4-5. (in Arabic). Khalaf, Norman Ali (1983). The Pine Bunting in Palestine. Gazelle: The Palestinian Biological Bulletin. ISSN 0178 – 6288. First Year. Number 1. July 1983. pp. 10-13. Al Gazelle : The Palestinian Biological Bulletin – Number 181 – January 2020
9
Salimiah, State of Kuwait. (In Arabic). http://issuu.com/drnormanalibassamkhalaf/docs/the_pine_bunting_in_palestine_gazel/1 Khalaf, Norman Ali Bassam (1984). The Palestine Bulbul. Gazelle: The Palestinian Biological Bulletin. Second Year. Number 5. May 1984. pp. 19-20. Hanweiler, Saarland, Federal Republic of Germany. (in Arabic). Khalaf, Norman Ali Bassam (1984-1985). The Weasel Project: Scientific Research on captive weasels (Mustela nivalis, Linnaeus 1766) in the Department of Zoology, University of Durham, Durham, England, during the Academic Year 1984-1985. Supervisor: Dr. Nigel Dunstone. Unpublished scientific research and data & scientific diary. Research Notebook. pp. 1-52. Khalaf, Norman Ali Bassam (1985). The Arabian Ostrich. Gazelle: The Palestinian Biological Bulletin. Department of Zoology, University of Durham, Durham, United Kingdom. Volume 3. Number 6. April 1985. pp. 1-7. (in Arabic). Khalaf, Norman Ali Bassam (1986). A List of the Birds of Palestine. Gazelle: The Palestinian Biological Bulletin. ISSN 0178-6288.Department of Zoology, University of Durham, Durham, United Kingdom. Number 8. Fourth Year. January 1986 AD, Jamada Oula 1406 H. pp. 1-24. (In Arabic and English). Khalaf, Norman Ali Bassam (1987). A Trip to Kuwait Zoo, State of Kuwait. Gazelle: The Palestinian Biological Bulletin. Rilchingen-Hanweiler, Federal Republic of Germany. Fifth Year, Number 13, Ramadan 1407 AH, April 1987 AD. pp. 1-5. (in Arabic). Khalaf-von Jaffa, Norman Ali Bassam (1991). A Trip to Zoo Budapest, Hungary. Gazelle: The Palestinian Biological Bulletin. Bonn-Bad Godesberg, Federal Republic of Germany. Number 21, Ninth Year, January 1991. pp. 1-4. Khalaf-von Jaffa, Norman Ali Bassam (1992). An Introduction to the Animal Life in Palestine. Gazelle. Gazelle: The Palestinian Biological Bulletin. Bonn-Bad Godesberg, Federal Republic of Germany. Number 30, Tenth Year, October 1992. pp. 1-7. (in Arabic). Khalaf-von Jaffa, Norman Ali Bassam (1994). An Introduction to the Animal Life in Palestine. Shqae'q Al-Nouma'n (Anemone coronaria). A Quarterly Magazine Issued by the Program EAI (Education for Awareness and for Involvement). Environmental Education / Children for Nature Protection. In Cooperation with Dept. of General and Higher Education. P.L.O., Palestine. Number 4.Huzairan (June) 1994.pp. 16-21. (in Arabic). Khalaf, Norman Ali Bassam (2001). The Extinct and Endangered Animals in Palestine. In: Gazelle: The Palestinian Biological Bulletin Home Page. Extinct and Endangered Animals and Reintroduction. http://gazelle.8m.net/photo3.html Khalaf-von Jaffa, Norman Ali (2004). Gazelle: Das Palästinensische Biologische Bulletin. Eine Wissenschaftliche Reise in Palästina, Arabien und Europa zwischen 1983 – 2004 / Gazelle: The Palestinian Biological Bulletin. A Scientific Journey in Palestine, Arabia and Europe between 1983 – 2004. ISBN 3-00-014121-9.Erste Auflage, Juli 2004: 452 Seiten. Zweite erweiterte Auflage, August 2004: 460 Seiten. Norman Ali Khalaf, BonnBad Godesberg, Germany. http://dr-norman-ali-khalaf-books.webs.com/ & eBook: https://www.yumpu.com/en/document/view/59405994/book-gazelle-theGazelle : The Palestinian Biological Bulletin – Number 181 – January 2020
10
palestinian-biological-bulletin-a-scientific-journey-in-palestine-arabia-and-europebetween-1983-2004-by-norman-ali-khalaf-von-jaffa-2004 Khalaf-von Jaffa, Norman Ali (2005). Aquatica Arabica. An Aquatic Scientific Journey in Palestine, Arabia and Europe between 1980 - 2005 / Aquatica Arabica. Eine Aquatische Wissenschaftliche Reise in Palaestina, Arabien und Europa zwischen 1980 - 2005. ISBN 3-00-014835-3. Erste Auflage, August 2005: 376 Seiten. Publisher: Norman Ali Khalaf, Rilchingen-Hanweiler, Bundesrepublik Deutschland & Sharjah, United Arab Emirates. http://dr-norman-ali-khalaf-books.webs.com/aquaticaarabica.htm & eBook: https://www.yumpu.com/en/document/view/59407769/book-aquatica-arabica-anaquatic-scientific-journey-in-palestine-arabia-and-europe-between-1980-2005-bynorman-ali-khalaf-von-jaffa-2005 & eBook: https://joom.ag/nN6L Khalaf-von Jaffa, Norman Ali (2005). The Rafah Zoo in the Rafah Refugee Camp, Gaza Strip, Palestine : A Story of Destruction by the Israeli Occupation Army. Gazelle: The Palestinian Biological Bulletin. Number 46, Twenty-third Year, October 2005, Ramadan 1426. pp. 1-11. Sharjah, United Arab Emirates. (in Arabic). Khalaf-von Jaffa, Norman Ali Bassam (2005). The Qalqilia Zoo and the Natural History Museum in the City of Qalqilia, West Bank, Occupied Palestine. Gazelle: The Palestinian Biological Bulletin. Number 47, Twenty-third Year, November 2005, Shawal 1426. pp. 1-10. Sharjah, United Arab Emirates. (in Arabic). Khalaf-von Jaffa, Norman Ali Bassam (Member of PALESTA) (2005). Palestinian Scientists and Technologists Abroad (PALESTA). Gazelle: The Palestinian Biological Bulletin. Number 47, Twenty-third Year, November 2005, Shawal 1426. pp. 11-12. Sharjah, United Arab Emirates (in Arabic). Khalaf, Norman Ali (2005, 2006, 2007). Chapter 3: Geography, Flora and Fauna. Pages 32-39. In: Palestine: A Guide. By Mariam Shahin, Photography by George Azar. CoAuthor: Norman Ali Khalaf. Northampton, Massachusetts: Interlink Publishing Group, 2005, 2006, 2007. xi + 471 pages. Appendices to page 500. http://ipsnewsite.mysite4now.com/journals.aspx?id=7323&jid=1&href=fulltext Khalaf-Sakerfalke von Jaffa, Norman Ali Bassam Ali Taher (Gründer) (seit Juni 2007). Yahoo! Deutschland Group: Fauna Palaestina. http://de.groups.yahoo.com/group/Fauna_Palaestina/ Khalaf-Sakerfalke von Jaffa, Norman Ali Bassam Ali Taher (2007). Felidae Arabica. A Zoological Journey in Palestine, Arabia and Europe between 1980-2007 / Felidae Arabica. Eine Zoologische Reise in Palästina, Arabien und Europa zwischen 1980-2007. ISBN 978-3-00-019568-6. Erste Auflage (First Edition), Juli (July) 2007, 300 pp. SelfPublisher: Norman Ali Khalaf, Rilchingen-Hanweiler, Deutschland & Sharjah, United Arab Emirates. (in Arabic, German and English). Doctoral Dissertation. Ashwood University, USA. Doctor of Science Degree in Zoology (Summa Cumm Laude) on 26.09.2007. http://dr-norman-ali-khalaf-books.webs.com/felidaearabica.htm & Doctoral Dissertation eBook : https://www.yumpu.com/en/document/view/59397999/doctorate-dissertationfelidae-arabica-by-norman-ali-bassam-khalaf-doctor-of-science-ashwood-universityusa-2007 Gazelle : The Palestinian Biological Bulletin – Number 181 – January 2020
11
Khalaf-Sakerfalke von Jaffa, Norman Ali Bassam Ali Taher (Gründer) (seit September 2007). Yahoo! Deutschland Group: Fauna Arabica. http://de.groups.yahoo.com/group/Fauna_Arabica/ Khalaf-Sakerfalke von Jaffa, Norman Ali Bassam Ali Taher (2007). Haywanat Filistin ( حيوانات فلسطينThe Animals of Palestine). Wikipedia, Al-Mawsu'aAl-Hurra (The Free Encyclopedia). Gazelle: The Palestinian Biological Bulletin. ISSN 0178-6288. Number 69, September 2007. Pp. 1-4. Sharjah, United Arab Emirates. http://ar.wikipedia.org/wiki/%D8%AD%D9%8A%D9%88%D8%A7%D9%86%D8%A7 %D8% AA_%D9%81%D9%84%D8%B3%D8%B7%D9%8A%D9%86 Khalaf-Sakerfalke von Jaffa, Dr. Sc. Norman Ali Bassam Ali Taher (2009). Flora and Fauna in Palestine. Gazelle: The Palestinian Biological Bulletin. Number 91, July 2009, Rajab 1430 AH. pp. 1-31. Sharjah, United Arab Emirates. http://flora-faunapalestine.webs.com/ Khalaf-von Jaffa, Dr. Norman Ali Bassam (2009). Fauna Palaestina – Part One. A Zoological Journey in Palestine, Arabia and Europe between 1983 – 2006 / Fauna Palaestina – Teil Eins. Eine Zoologische Reise in Palästina, Arabien und Europa zwischen 1983 – 2006. ISBN 978-9948-03-865-8. Erste Auflage/First Edition, September 2009: 412 Seiten/Pages. Self Publisher: Dr. Norman Ali Bassam Khalaf-von Jaffa, Sharjah, United Arab Emirates & Rilchingen-Hanweiler, Bundesrepublik Deutschland. http://dr-norman-ali-khalaf-books.webs.com/faunapalaestinapart1.htm & eBook: https://www.yumpu.com/xx/document/view/59498633/fauna-palaestina-1-bookby-dr-norman-ali-khalaf-2009 Dr. Sc. Norman Ali Bassam Ali Taher Khalaf-Sakerfalke von Jaffa, Zoologist, Ecologist and Geologist: The Scientific References (1980-2009). http://dr-norman-ali-khalafreferences.webs.com/ Khalaf-Sakerfalke von Jaffa, Dr. Sc. Norman Ali Bassam Ali Taher (2009). An ancient Arabian Ostrich (Struthio camelus syriacus) egg-shell from the Village of Qidfa, Emirate of Fujairah, United Arab Emirates. Gazelle: The Palestinian Biological Bulletin. ISSN 0178 – 6288. Twenty-seventh Year, Number 96, December 2009, Thu Al Hijja 1430 AH. pp. 1-25. Sharjah, United Arab Emirates. http://arabian-ostrich.webs.com/ Khalaf-Sakerfalke von Jaffa, Dr. Sc. Norman Ali Bassam Ali Taher (2009). The RedNecked Ostrich (Struthio camelus camelus Linnaeus, 1758) at Dubai Zoo, Dubai, United Arab Emirates. Gazelle: The Palestinian Biological Bulletin. ISSN 0178 – 6288. Twentyseventh Year, Number 96, December 2009, Thu Al Hijja 1430 AH. pp. 26-28. Sharjah, United Arab Emirates. http://arabian-ostrich.webs.com/ostrichdubaizoo.htm Khalaf-Sakerfalke von Jaffa, Dr. Sc. Norman Ali Bassam Ali Taher (Photographer) (09 December 2009). Photo: Ancient Arabian Ostrich Egg Shell / Alte arabische Straußeneischale. Emirat Fujairah, V. A. Emirate. Fotocommunity. www.fotocommunity.de/pc/pc/mypics/1213259/display/22065236 Khalaf-Sakerfalke von Jaffa, Dr. Sc. Norman Ali Bassam Ali Taher (Photographer) (15 December 2009). Photo: Ancient Arabian Ostrich Egg Shell, Qidfa Village, Emirate of Fujairah, United Arab Emirates. Gazelle : The Palestinian Biological Bulletin – Number 181 – January 2020
12
www.fotocommunity.de/pc/pc/mypics/1213259/display/21831255 Khalaf-Sakerfalke von Jaffa, Dr. Sc. Norman Ali Bassam Ali Taher (Fotograf) (04 August 2010). Foto: Nordafrikanische Strauß, Dubai Zoo, Dubai, Vereinigte Arabische Emirate. Fotocommunity. www.fotocommunity.de/pc/pc/mypics/1213259/display/21993963 Khalaf-von Jaffa, Dr. Sc. Norman Ali Bassam Ali Taher (2010). Fauna Emiratus-Part One. Zoological Studies in the United Arab Emirates between 2004 - 2009. /Fauna Emiratus – Teil Eins. Zoologische Studien in die Vereinigten Arabischen Emirate zwischen 2004 - 2009. ISBN 978-9948-15-462-4. Erste Auflage/First Edition, November 2010: 350 Seiten / Pages. Self Publisher: Dr. Norman Ali Bassam Khalaf-von Jaffa, Dubai and Sharjah, United Arab Emirates & Rilchingen-Hanweiler, Bundesrepublik Deutschland. http://dr-norman-ali-khalaf-books.webs.com/faunaemiratuspart1.htm & eBook: https://www.yumpu.com/xx/document/view/59546804/fauna-emiratus-part-1zoological-studies-in-the-united-arab-emirates-between-2004-2009-by-dr-norman-alibassam-khalaf-von-jaffa-2010 Khalaf-von Jaffa, Dr. Norman Ali Bassam (2012). Fauna Palaestina – Part Two. Zoological Studies in Palestine between 1983 – 2009/ Fauna Palaestina – Teil Zwei. Zoologische Studien in Palästina zwischen 1983 – 2009. ISBN 978-9948-16-667-2. 1. Auflage / First Edition : July 2012, Shaaban 1433 H. 208 Seiten / Pages (Arabic Part 120 Pages and the English Part 88 Pages). Publisher: Dar Al Jundi Publishing House, Jerusalem, Palestine. http://dr-norman-ali-khalafbooks.webs.com/faunapalaestinapart2.htm & & eBook: https://www.yumpu.com/en/document/view/59602956/fauna-palaestina-part-2book-by-dr-norman-ali-khalaf-von-jaffa-2012 Khalaf-von Jaffa, Dr. Norman Ali Bassam (2013). Fauna Palaestina – Part Three. Zoological Studies in Palestine between 2005 – 2012 / Fauna Palaestina – Teil Drei. Zoologische Studien in Palästina zwischen 2005 – 2012.ISBN 978-9950-383-35-7. Erste Auflage / First Edition : July 2013, Shaaban 1434 H. 364 pages (English Part 350 Pages and the Arabic Part 14 Pages). Publisher: Dar Al Jundi Publishing House, Jerusalem, State of Palestine. http://dr-norman-ali-khalafbooks.webs.com/faunapalaestinapart3.htm Khalaf-von Jaffa, Prof. Dr. Norman Ali Bassam (2014). Fauna Palaestina – Part Four. Zoological Studies in Palestine between 1983 – 2014 / Fauna Palaestina – Teil Vier. Zoologische Studien in Palästina zwischen 1983 – 2014. ISBN 978-9950-383-77-7. Erste Auflage / First Edition : July 2014, Ramadan 1435 H. pp. 456 (English part 378 pages and Arabic part 78 pages). Publisher: Dar Al Jundi Publishing House, Al-Quds (Jerusalem), State of Palestine. http://fauna-palaestina-part1.webs.com/faunapalaestina4.htm Khalaf-Sakerfalke von Jaffa, Prof. Dr. Sc. Norman Ali Bassam Ali Taher (2014). The Animals living inside and around the Sacred Mosque (Al-Masjid Al-Haram) in Makkah Al-Mukarramah, Kingdom of Saudi Arabia. Gazelle: The Palestinian Biological Bulletin. ISSN 0178 – 6288. Number 116, August 2014, Shawal 1435 AH. pp. 1-23. Dubai and Sharjah, United Arab Emirates. http://animals-of-makkah.webs.com/ Gazelle : The Palestinian Biological Bulletin – Number 181 – January 2020
13
Khalaf-Sakerfalke von Jaffa, Prof. Dr. Sc. Norman Ali Bassam Ali Taher (2014). Dinasurat Al Quds (Dinosaurs of Jerusalem). Dialogue with Prof. Dr. Norman Khalafvon Jaffa. National Geographic Arabiya Magazine (October 2014, Pages 52--53). https://www.flickr.com/photos/50022881@N00/15412670595 Khalaf-von Jaffa, Prof. Dr. Norman Ali Bassam Ali Taher (2015). Dr. Norman Ali Khalaf Award for Biodiversity in Palestine 2012 / 2013 : Essay and Photography Contest of the Palestine Sunbird (Cinnyris osea Bonaparte, 1856). Gazelle: The Palestinian Biological Bulletin. ISSN 0178 – 6288. Number 124, April 2015. pp. 1-21. Dubai and Sharjah, United Arab Emirates. http://dr-norman-ali-khalafreferences.webs.com/drnormanakhalafaward.htm Khalaf-Sakerfalke von Jaffa, Prof. Dr. Norman Ali Bassam Ali Taher (2015). Plants and Animals unique to Palestine. Gazelle: The Palestinian Biological Bulletin. ISSN 0178 – 6288. Number 125, May 2015. pp. 1-18. Dubai and Sharjah, United Arab Emirates. http://flora-fauna-palestine-2.webs.com/ Khalaf-von Jaffa, Prof. Dr. Norman Ali Bassam (2015). Fauna Palaestina – Part Five. Zoological Studies in Palestine between 1983 – 2016 / Fauna Palaestina – Teil Fünf. Zoologische Studien in Palästina zwischen 1983 – 2016. ISBN 978-9950-383-92-0. Erste Auflage / First Edition : July 2015, Ramadan 1436 H. 448 pp. (English Part 304 Pages and the Arabic Part 144 Pages). Publisher: Dar Al Jundi Publishing House, Al-Quds (Jerusalem), State of Palestine. http://fauna-palaestina-books.webs.com/ Khalaf-Sakerfalke von Jaffa, Prof. Dr. Norman Ali Bassam Ali Taher (2016). The First Palestinian Wildlife Photography Exhibition in the Gaza Strip in November 2014. Gazelle: The Palestinian Biological Bulletin. ISSN 0178 – 6288. Number 138, June 2016. pp. 1-35. Dubai and Sharjah, United Arab Emirates. http://dr-norman-ali-khalafreferences.webs.com/gazawildlifephoto2014.htm Khalaf-Sakerfalke von Jaffa, Prof. Dr. Sc. Norman Ali Bassam Ali Taher (2016). The Presence of the Hoopoe (Upupa epops Linnaeus, 1758) in the Gardens of Sharjah, United Arab Emirates. Gazelle: The Palestinian Biological Bulletin. ISSN 0178 – 6288. Number 139, July 2016. pp. 1-23. Dubai and Sharjah, United Arab Emirates. http://animals-ofuae.webs.com/ Khalaf-Sakerfalke von Jaffa, Prof. Dr. Sc. Norman Ali Bassam Ali Taher (2016). The Hoopoe )Upupa epops Linnaeus, 1758) in Palestine. Gazelle: The Palestinian Biological Bulletin. ISSN 0178 – 6288. Number 140, August 2016. pp. 1-21. Dubai and Sharjah, United Arab Emirates. http://animals-of-palestine-2.webs.com/ Khalaf-Sakerfalke von Jaffa, Prof. Dr. Sc. Norman Ali Bassam Ali Taher (2016). The Desert Tawny Owl (Strix hadorami Kirwan, Schweizer and Copete, 2015): New Species of Bird Discovered in Palestine. Gazelle: The Palestinian Biological Bulletin. ISSN 0178 – 6288. Number 142, October 2016. pp. 1-13. Dubai and Sharjah, United Arab Emirates. http://birds-of-palestine.webs.com/desert-tawny-owl Khalaf-Sakerfalke von Jaffa, Prof. Dr. Sc. Norman Ali Bassam Ali Taher (2016). The Pine Bunting (Emberiza leucocephalos Gmelin, S.G., 1771) in Palestine. Gazelle: The Palestinian Biological Bulletin. ISSN 0178 – 6288. Number 143, November 2016, pp. 1-6. Dubai and Sharjah, United Arab Emirates (In Arabic). http://birds-of-palestine.webs.com/pineGazelle : The Palestinian Biological Bulletin – Number 181 – January 2020
14
bunting Khalaf-Sakerfalke von Jaffa, Prof. Dr. Sc. Norman Ali Bassam Ali Taher (2016). Haywanat Falastin (Fauna of Palestine) حيوانات فلسطين. Gazelle: The Palestinian Biological Bulletin. ISSN 0178 – 6288. Number 144, December 2016, pp. 1-18. Dubai and Sharjah, United Arab Emirates (In Arabic). http://animals-of-palestine-2.webs.com/fauna-ofpalestine-arabic Khalaf-Sakerfalke von Jaffa, Prof. Dr. Sc. Norman Ali Bassam Ali Taher (2017). Bateleur Eagle (Terathopius ecaudatus Daudin, 1800): A Rare Record in Palestine in 2015 /Der Gaukler (Terathopius ecaudatus) in Palästina / من الطيور النادرة في: العُقاب البهلواني أو العُقاب ال ُمصفق فلسطين. Gazelle: The Palestinian Biological Bulletin. ISSN 0178 – 6288. Number 145, January 2017. pp. 1-22. Dubai and Sharjah, United Arab Emirates. (In English, German and Arabic). http://birds-of-palestine.webs.com/bateleur-eagle Khalaf-Sakerfalke von Jaffa, Prof. Dr. Sc. Norman Ali Bassam Ali Taher & Nora Norman Ali Khalaf (Photography) (2017). Greater Flamingos (Phoenicopterus roseus Pallas, 1811) at Ras Al Khor Wildlife Sanctuary in Dubai, United Arab Emirates. Gazelle: The Palestinian Biological Bulletin. ISSN 0178 – 6288. Number 146, February 2017. pp. 1-25. Dubai & Sharjah, United Arab Emirates. http://animals-of-uae.webs.com/greaterflamingo Khalaf-Sakerfalke von Jaffa, Prof. Dr. Sc. Norman Ali Bassam Ali Taher (2017). Melanistic Greater Flamingo (Phoenicopterus roseus Pallas, 1811) Records from Palestine and Cyprus Between 2013-2015. Gazelle: The Palestinian Biological Bulletin. ISSN 0178 – 6288. Number 147, March 2017. pp. 1-16. Dubai and Sharjah, United Arab Emirates. http://birds-of-palestine.webs.com/ Khalaf-Sakerfalke von Jaffa, Prof. Dr. Sc. Norman Ali Bassam Ali Taher (2017). The Senegal Thick-Knee or Stone-Curlew (Burhinus senegalensis Swainson, 1837): New Record for Palestine. Gazelle: The Palestinian Biological Bulletin. ISSN 0178 – 6288. Number 148, April 2017. pp. 1-13. Dubai and Sharjah, United Arab Emirates. http://animals-of-palestine-2.webs.com/senegal-thick-knee Khalaf, Sharif Prof. Dr. Sc. Norman Ali Bassam Ali Taher Mohammad Ahmad Ahmad Mostafa Abdallah Mohammad (Abu Nora) (2018). The Family of Sharif Hajji Taher Mohammad Ahmad Ahmad Mostafa Khalaf (Abu Othman). A Pictorial History Book of a Palestinian Family from Jaffa in the Twentieth Century. ISBN 978-9950-974-40-1. First Edition, October 2018, Safar 1440 Hijri. 120 pp. In Arabic. Publisher: Prof. Dr. Norman Khalaf Department for Environmental Research and media, National Research Center, University of Palestine, Gaza, State of Palestine. https://family-taherkhalaf.webs.com/ & eBook: https://www.yumpu.com/xx/document/view/62242473/book-family-taher-khalaf2018 Khalaf-Prinz Sakerfalke von Jaffa, Sharif Prof. Dr. Sc. Norman Ali Bassam Ali Taher Mohammad Ahmad Ahmad Mostafa Abdallah Mohammad (January 2020). A Rare Stamp from Palestine: The Rockhopper Penguin (Eudyptes chrysocome) & Gentoo Penguin (Pygoscelis papua) Stamp. Gazelle: The Palestinian Biological Bulletin. ISSN 0178 – 6288. Volume 38, Number 181, January 2020, pp. 1-15. Published by Prof. Dr. Norman Gazelle : The Palestinian Biological Bulletin – Number 181 – January 2020
15
Ali Khalaf Department for Environmental Research and Media, National Research Center, University of Palestine, Gaza, State of Palestine. https://palestinestamps.webs.com/
Khalaf, Ola Mostafa (Fotografin) (27 July 2007). Foto: Strauß, Qaryet Al Asad (Löwe Dorf), Kairo-Alexandria Wüsten-Straße, Ägypten. Fotocommunity. www.fotocommunity.de/pc/pc/extra/buddies/display/21947580 Khalaf, Ola Mostafa (Fotografin) (02 August 2010). Foto: Afrikanische Straußeneier / African Ostrich Egg-shell. Souk Al Arsah, Sharjah, Vereinigte Arabische Emirate. Fotocommunity. www.fotocommunity.de/pc/pc/extra/buddies/display/22032858 Wikipedia. Eastern rockhopper penguin. https://en.wikipedia.org/wiki/Eastern_rockhopper_penguin Wikipedia. Gentoo Penguin. https://en.wikipedia.org/wiki/Gentoo_penguin Wikipedia. Northern rockhopper penguin. https://en.wikipedia.org/wiki/Northern_rockhopper_penguin Wikipedia. Rockhopper Penguin. https://en.wikipedia.org/wiki/Rockhopper_penguin Wikipedia. Southern Rockhopper Penguin. https://en.wikipedia.org/wiki/Southern_rockhopper_penguin Wood, Ian (24 January 2009). Photo: A Gentoo Penguin (Pygoscelis papua) parent feeding chick. The Internet Bird Collection (IBC). https://www.hbw.com/ibc/photo/gentoopenguin-pygoscelis-papua/parent-feeding-chick
Gazelle : The Palestinian Biological Bulletin – Number 181 – January 2020