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biology

THE CORE

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Library of Congress Cataloging-in-Publication Data

Names: Simon, Eric J. (Eric Jeffrey), 1967- author.

Title: Biology : the core / Eric J. Simon.

Description: Third edition. | Hoboken : Pearson, [2020] | Includes index.

Identifiers: LCCN 2018024849| ISBN 9780134891514 | ISBN 0134891511

Subjects: LCSH: Biology.

Classification: LCC QH308.2 .S56 2020 | DDC 570–dc23

LC record available at https://lccn.loc.gov/2018024849

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ISBN 10: 0-134-8915-11;

ISBN 13: 978-0-134-89151-4 (Student edition)

ISBN 10: 0-135-27165-7;

ISBN 13: 978-0-135-27165-0 (Looseleaf Edition)

ABOUT THE AUTHOR

Eric J. Simon is a professor in the Department of Biology and Health Science at New England College in Henniker, New Hampshire. He teaches introductory biology and human biology to non-science majors, as well as a tropical marine biology course that introduces non-science-majors to underwater field work in Belize. Dr. Simon received a B.A. in biology and computer science and an M.A. in biology from Wesleyan University and a Ph.D. in biochemistry from Harvard University. His research focuses on innovative ways to use technology to improve the teaching and learning of science, particularly to non-science majors. He lives in rural New Hampshire with his wife, two boys, a rotating set of Cavalier King Charles Spaniels, a few dozen chickens, and a leopard gecko. Dr. Simon is the lead author of the introductory non-majors biology textbooks Campbell Essential Biology (7th ed.) and Campbell Essential Biology with Physiology (6th ed.), and a co-author of the introductory biology textbook Campbell Biology: Concepts and Connections (9th ed.), all published by Pearson Benjamin Cummings.

“I dedicate this book to my inspirational partners at Benjamin Cummings and Pearson Education, including (in order of appearance) Beth Wilbur, Frank Ruggirello, Chalon Bridges, Ginnie Simione-Jutson, Josh Frost, Lauren Harp, Evelyn Dahlgren, and Alison Rodal. Thank you all for providing me with tremendous support and continuous inspiration to improve biology education.”

PREFACE

To the Student,

Being a college student today means juggling many priorities: work, school, extracurricular activities, family. If you’re reading this book, you’ve probably enrolled in your first college science course, and it may be the only one you’ll ever take. With so many priorities competing for your attention, you may be unsure how to fit studying biology into your busy life. Good news: This book is written specifically for you!

Over the years, I’ve seen students in my classes striving to succeed while also wishing to be as productive as possible with their study time. Biology: The Core was designed from the ground up to help you learn efficiently and thrive in this course. Only the most important and relevant information—the core of biology content—is included. These biological concepts are displayed in highly visual, consistent, and approachable two-page modules that guide you along a clear learning path, so that reading your textbook is more a pleasure than a chore.

You might also be wondering how this course—and biology in general—applies to your own life. Luckily, this is easy to address, since issues like nutrition, cancer, vaccines, and genetically modified foods directly affect you and those you care about. For the Third Edition, new modules were added that address these and other current issues directly, so that you may better see how biology is relevant to your life. Other modules help you critically evaluate the scientific-sounding claims that constantly bombard you, and how to distinguish valid scientific claims from bogus ones.

The Biology: The Core textbook is paired with a robust online library, Mastering Biology, that contains videos, animations, current events, and interactive tutorials that help you draw connections between the course material and the world around you. Questions you might have about many topics will be addressed in this online complement to your textbook. It is filled with helpful multimedia tools that allow you to gain a thorough understanding of the content so that you can succeed in your course. References to Mastering Biology at the top of many modules point you to the most helpful online tools.

I hope that Biology: The Core meshes with your goals and your priorities, acting as a useful guide for this course and addressing questions you run into in your broader life. Please feel free to drop me an email to tell me about your experience with Biology: The Core or to provide feedback (good or bad!) regarding the text or online resources.

Best wishes for a successful semester—and enjoy the big adventure of biology! It’s not only in the pages of this book, but everywhere around you.

To the Instructor,

In a world with so many options for non-major biology textbooks, what makes this one different? The answer is: a focus on today’s students. We’ve all watched our non-science-major students struggle with the depth of material and relating biology to their lives. Which concepts do non-science students need to know in order to understand the relevance of biology? If we pare down the content and focus on the most important take-home lessons—the information that we hope students will remember 10 years after your course—what remains is the core: a set of essential biological concepts that presents the big picture, providing students with a scientific basis for the issues they will confront throughout their lives.

Biology: The Core is a different kind of textbook, one that presents information in small chunks using a nonlinear, engaging, visual style. The book contains only the most essential content for each topic. All information is presented in stand-alone two-page modules that fully integrate text and art into a single teaching tool. Modules can be read in any order and each module stands alone (without references to other modules), allowing you the flexibility to assign topics in whatever sequence best suits your course.

For the Third Edition of Biology: The Core, content was revised based on feedback received from professors and students using the text from around the country. The specific changes are detailed on the next page, but the overall approach is to ensure that the core content is approachable and clearly connected to students’ lives. Included in this new edition are a series of “Core Issue” modules. These ten special modules highlight current topics that your students may hear about frequently—vaccines, antibiotic resistance, diabetes, and cancer, to name a few—and tie them to the core content, showing students the relevancy of basic biology knowledge.

Over the last few years, in my communication with many instructors around the country, I’ve also observed that those of us teaching biology to non-majors are increasingly emphasizing science literacy skills above all else. For this reason, the Third Edition places particular emphasis on teaching students to think scientifically and helping them appreciate and apply critical thinking skills to their own lives. A largely rewritten Chapter 1 brings many of these important skills together.

The printed text is paired with Mastering Biology, an online tutorial platform that allows you to reinforce the book content and expand on the basic concepts presented in each module as needed. The activities and resources in Mastering Biology also offer you the flexibility to incorporate a wide variety of applications and current issues—including several ones new to this edition—into your teaching. Unlimited by the particular set of examples printed in a static textbook, a rich collection of online resources—including Current Topic PowerPoint presentations, news videos, Current Events news articles, and interactive tutorials—enables you to connect the core content to interesting, relevant, and timely issues that are important to you and your students. Forty—including 18 brand new, Guided Video Tours found within Mastering Biology are designed to help students learn to use the textbook and to hone their study skills.

I hope that the aims of Biology: The Core resonate with the teaching and learning goals of your non-major introductory biology course. Feel free to send an email telling me about your course and your students, to provide feedback regarding the text or the online resources, or just to chat about the non-major course in general—it’s my favorite topic of conversation!

Best wishes for a successful semester,

Biology: The Core, Third Edition, contains many helpful updates

The Third Edition of Biology: The Core was created in response to extensive feedback from professors and students. The goal of the new edition is to enhance teaching and learning for non-major students by increasing the relevancy of the material and placing a greater emphasis on science literacy skills.

CHAPTER 1 AN INTRODUCTION TO THE SCIENCE OF LIFE

For many students, this introductory biology course is their only exposure to college-level science. Many instructors believe that the most important goal of such a course is to communicate how science is conducted and how the process of science can be used to make important decisions. In the Third Edition of Biology: The Core, we dedicate an entire chapter to covering this vital topic, with modules that present the process of science in a more realistic and relevant context. Chapter 1, “An Introduction to the Science of Life,” promotes critical thinking and demystifies how science works. A new presentation of the process of science emphasizes the lack of formality in the process and how it proceeds in the real world.

Additional updated and new modules help students distinguish hypotheses from theories and explain the meaning of the word “fact”; discuss basic research methods; distinguish scientific thought from pseudoscience; and explain how to recognize reliable sources and the process of peer review. This chapter should help students obtain a clearer picture of how scientific thinking differs from other ways of viewing the world and how it can be applied in their own lives.

Module 1.4 presents the process of science as it actually occurs.

NEW EXAMPLES AND PEDAGOGICAL IMPROVEMENTS

In addition to those already mentioned, many changes in the book were implemented to increase accuracy and currency. For example, module 6.14 includes new information about the CRISPR-Cas9 gene editing system.

Other examples of content updates include new data on the links between obesity and cancer, changes to human population growth patterns, and new data on the reliability of various methods of contraception. Throughout the book, photos and art were improved and updated to make them more attractive and better able to convey the pedagogical points. Every module contains a “fun fact” intended to invoke a “That’s cool!” reaction from your students; many of these have been updated to be even more engaging.

CORE ISSUE MODULES

New to this edition are ten Core Issue modules. Each one presents a current relevant topic. This edition includes modules on nutrition, cancer, vaccines, athletic cheating, genetically modified organisms, agriculture, MRSA, climate change, biodiversity hot spots and diabetes. Each Core Issue module helps relate the basic biology content to the issue at hand.

In this example, the Core Issue: Nutrition module shows how the digestive system (Module 11.4) delivers small molecules (Modules 2.9–2.12) through the circulatory system (Module 11.9) to the mitochondria (Module 4.2).

Additionally, every Core Issue module includes a description of one or more relevant scientific studies. Data is presented, and the larger lessons of that study are discussed in relation to the material presented in Chapter 1. For example, the Core Issue: Biodiversity Hot Spots module introduces a study showing the relationship between forest fragmentation in the northeastern United States and the prevalence of mosquitoes that carry Lyme disease. For each Core Issue module, the chosen study illustrates one or more science skill presented in Chapter 1. In this example, students are led to understand independent and dependent variables, as originally discussed in module 1.6.

NEW MULTIMEDIA

The Biology: The Core textbook is designed to pair with the online resources (videos, animations, current issues, practice assignments, and more) available in Mastering Biology. New references within many modules point students toward the most helpful multimedia supplements. This edition of Biology: The Core includes 18 new Guided Video Tours, for a total of 40 videos that walk students through modules, teaching them how to organize their studying. These videos can be particularly helpful to help students working on their own (as part of flipped classrooms or distance learning, for example).

CORE IDEA

In response to feedback, the Third Edition of Biology: The Core improves teaching and learning by making the material clearer and more relevant to non-science-major students.

Mastering Biology

WATCH the Video Tutor Session for this module

Icons placed at the top of modules remind students to supplement their learning with online multimedia.

How are the Core Issue modules different from every other module in The Core?

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Help students see biology’s relevance by focusing on core concepts

Biology: The Core presents essential biological concepts, using a visual and hybrid approach. The 12-chapter textbook uses dynamic illustrations organized into concise, self-contained two-page modules that focus students’ attention on what is most relevant. The text pairs with Mastering Biology to offer flexible assignment options and extensive support materials that allow instructors to tailor the content to the way they teach and maximize student engagement.

Build your course around . . .

Each core biological concept is presented as a two-page module that can stand on its own and be read in any order. Each module in the text contains only the most essential content for any concept. The efficient organization of each module helps students focus their attention on key information and guides them through—from the clearly stated concept at the start to the “core question” that checks their understanding at the end.

relevant and current high interest topics

NEW! Core Issue Modules highlight relevant current issues like vaccinations, antibiotic resistance, cancer, and more. Each new module relates the core issue to biological concepts to help students see the relevancy of the course material, as well as connections across chapter concepts.

Each Core Issue Module is supported by a prebuilt assignment in Mastering Biology, while a Topic Guide and a Current Topic Lecture presentation in the Instructor Resources area of Mastering Biology helps instructors bring the issue into the classroom.

Develop students’ scientific literacy skills

Evaluating Science in the Media Coaching Activities guide students through a step-by-step process for evaluating the authority, motivation, and reliability of online sources of scientific information. Topics include genetically modified organisms, head injuries, tanning and skin cancer, and more.

GraphIt!

Coaching Activities help students read, interpret, and create graphs that explore real environmental issues using real data. All 10 activities explore current topics such as the carbon footprint of food, fresh water availability, and ocean acidification in an entirely new mobile experience with accessible design.

Engage students with active learning

NEW! Ready-to-Go Teaching Modules make use of teaching tools for before, during, and after class, including new ideas for in-class activities. Each of the 10 modules for The Core highlights a specific current issue and suggests how to incorporate Mastering Biology, active learning resources including Learning Catalytics, and instructor resources. These modules can be accessed through the Instructor Resources area of Mastering Biology.

UPDATED! Guided Reading Activities are organized around each module and provide students with basic questions that guide them through the module, using an active reading approach. The worksheets offer an easy, low-tech way to assign work outside of or during class as a group work activity. These are available in the Mastering Biology Study Area, in the Instructor Resources, and in Pearson Collections.

Reach every student with Pearson eText and Dynamic Study Modules

Dynamic Study Modules help students study effectively—and at their own pace. Each module poses a series of questions about a course topic, which adapt to each student’s performance and offer personalized, targeted feedback to help them master key concepts.

Students can easily review their answers and monitor their own progress and understanding of key concepts as they move through each module.

Pearson eText is a simpleto-use, mobile optimized, personalized reading experience available within Mastering. It allows students to easily highlight, take notes, and review key vocabulary all in one place—even when offline. Integrated videos engage students and give them help when they need it. Pearson eText is available within Mastering Biology when it comes with a new book; students can also purchase Mastering with Pearson eText online. For instructors not using Mastering, Pearson eText can also be adopted on its own as the main course material.

Deliver trusted content in Mastering Biology

NEW! Topic Overview videos, created by the author, introduce key concepts and vocabulary. These brief, engaging videos introduce topics that will be explored in greater depth in class.

Topics include:

• Macromolecules

• Ecological Organization

• Mechanisms of Evolution

• An Introduction to Structure and Function

• Interactions Between the Respiratory and Circulatory Systems

• DNA Structure and Function

• And more!

NEW and UPDATED! Guided Video Tours, developed and narrated by author Eric Simon, present a brief “minilecture” that walks students through key concepts and module content presented in the text. All Guided Video Tours can be assigned as a coaching activity with personalized feedback in Mastering Biology, and are also embedded in the eText.

Resources to help you build your class, your way

Biology: The Core includes a full suite of instructor support materials in the Instructor Resources area in Mastering Biology to help you build your course. Resources include Ready-to-Go Teaching Modules; current topic lecture slides and topic guides that help you integrate biology content with relevant current topics; an active learning resource guide; animations, videos, and lecture presentations to show in class; all images from the text; a testbank; and free access to our unique Instructor Exchange website, where you can share ideas with other non-majors biology faculty.

ACKNOWLEDGMENTS

There are many people to thank and acknowledge for their contributions to the Third Edition of Biology: The Core. One of the most gratifying aspects of writing a book is the opportunity to interact with so many skilled professionals at Pearson Education and so many talented biology colleagues from around the country. While any problems or mistakes within this book or any of the materials related to it are solely my responsibility, the successful aspects are due to the efforts of many people.

A few people deserve special mention for their deep contributions; these people form the core of The Core. Alison Rodal, as Courseware Portfolio Manager, worked with me and the rest of the team nearly every day, bringing a broad array of talents to bear on this project. Alison’s consistently positive energy and her diplomatic skills were invaluable at several key junctures in the book’s development. Alison reminds me every day that our overriding goal is to produce a book that will help students and teachers, and her guidance is invaluable. I will always owe a deep debt of gratitude to Beth Wilbur, Director of Portfolio Management, whose vision, understanding, and unwavering high standards have benefited every project since she brought me into the Pearson family over a decade ago. Courseware Editorial Assistants Sherry Wang, Summer Giles and Chelsea Noack were a continuous help in coordinating reviews and solving countless problems, always with a buoying positive energy. Ginnie Simione Jutson, Courseware Director of Content Development, provided her masterful oversight of all matters editorial and gave invaluable help when it was most needed.

I love my editors! All members of the Pearson editorial team bring decades of experience and talent to every project they take on. Senior Developmental Editor Evelyn Dahlgren has been my rock for this edition, building on the work and talents of Mary Ann Murray in the Second Edition. Evelyn’s expertise with both words and art were of great help in preparing this edition. Hilair Chism used her keen sense of art development to design and improve many of the new features found in this edition. You can see Hilair’s unique skills shine in Chapter 1 and in the Core Issue modules, both of which glimmer with her talent. Content Producer Ami Sampat proved to be a wise, capable, and gentle guide through the seemingly countless deadlines that comprise the writing of a new edition. Thank you to Indexer Robert Swanson for making the book much more user-friendly (see “Indexer, continued gratitude for”). Design Manager Mark Ong for his work on the Core Issue modules and designer Tani Hasegawa refreshed the design for this third edition and also designed the cover. Once the text was written and the art developed, the production team took over, turning countless individual ideas into the reality of the book you see before you. For this, we thank Photo Researcher John Paul Belciña of Wanderlust Photos, Pearson Rights & Permissions Project Manager Matt Perry, Proofreaders Pete Shanks and Marleis Roberts, copyeditor Julie Kennedy, and Project Manager Margaret McConnell of Integra, which also provided expert composition services. Mark Mykytiuk and his team at Imagineering lent their talents to art development and art creation.

I am deeply indebted to the many talented thinkers who were kind enough to share their expertise. Jim Newcomb of New England College provided insight for the revision of several chapters and always happily fielded my questions at any time no matter how goofy or obscure. I am also very grateful to Amanda Marsh who provided both proofreading acumen and science knowledge as she read every chapter of the book. Special thanks also go to Juliette Madan, M.D. (for help with prenatal screening), Tim Ritchie (for help understanding how cars work), Bart Zuckerman (for advice on legal affairs), Aaron Daniels of Curry College (for advice about matters philosophical), Elyse Carter Vosen of College of Saint Scholastica (for advice on multiculturalism), Andrew R. Swanson of State College of Florida – Manatee/Sarasota (who speaks for the protists), Lori Koziol of New England College (for information on biofilms), Helen Walter of Mills College (for providing underwater photos), Terry Austin of Temple College (for help with A&P best practices), Forest Simon (for providing photos), Ceilidh Kehoe and several students at New England College (for posing for photos), Tod Ramseyer and Mike Gardner of New England College (for providing equipment), Heidi Hammel of the Association of Universities for Research in Astronomy, Inc. (for help with water on Mars), Melissa Gallagher of The Grapevine Family & Community Resource Center (for providing advice about food assistance), and Ira Bragg-Grant of the American Sickle Cell Anemia Association. I’d also like to thank my colleagues at New England College—especially Jim Newcomb, Lori Koziol, Geoff Cook, Deb Dunlop, Mike Gardner, and Elizabeth Harper—for providing examples of student-centric and skills-based teaching that significantly informed Chapter 1 of this text. I am also indebted to the many faculty colleagues around the country who reviewed chapters of The Core, met with me, or provided feedback on their own classroom experiences. You can find a complete list of all those who have made contributions on pages xix–xxi; thanks go to every name on that list. I would also like to thank all of my colleagues at New England College— including Mark Mitch, Bryan Partridge, Wayne Lesperance, and Michelle Perkins—for providing support of various kinds during this project. Additionally, I would like to thank the nurses of the Ambulatory Care Center at Concord Hospital for providing excellent medical care while also helpfully facilitating my crazy work schedule.

Of equal importance to the book itself are the many supplements that accompany it. Many teaching colleagues provided help with writing supplements for The Core. In particular, I thank Test Bank author Wiline Pangle of Central Michigan University, Instructor’s Guide author Lori Nicholas of Northwest Vista College, PowerPoint presentation author Wendy Kuntz of Kapi’Olani Community College and University of Hawai’i, and Active Reading Guide author Dana Kurpius of Elgin Community College. Special thanks go to Tina Hopper of Missouri State University and Andrew Swanson of State College of Florida for writing the Reading Questions, Jennifer Wiatrowski of Pasco Hernando State College for writing the Practice Tests, and Angel Nickens of Northwest

Mississippi Community College for writing the Dynamic Study Modules. I thank Ami Sampat who expertly coordinated all of the instructor resources, which is no small task given their number and variety. Many people at Pearson also helped produce the media that accompanies this text. In particular, Rich Media Producer Mireille Pfeffer, Mastering Biology Development Editor Sarah Jensen, and Amanda Kaufman of Thrasher Editorial for her work on the videos that compliment this text. From the engineering side of Mastering Biology, I thank Katie Foley, Caroline Ross, and Ashley Gordon.

After a book is written, the marketing team steps in to ensure that the complete story is told to all who might benefit. For this we thank Product Marketing Manager Alysun Burns and Field Marketing Manager Kelly Galli. Acting as the final facilitators of the long journey from author to student,

I thank the entire Pearson Education sales team including all the sales managers, publisher’s representatives, and technology specialists who work tirelessly every semester to help students learn and instructors teach.

In closing, I beg forgiveness from those who lent their unique talents to this book but who I failed to mention—I hope you will excuse my oversight and know that you have earned my gratitude.

With deepest, sincerest, and humblest thanks and respect to all who contributed their talents to Biology: The Core, Third Edition,

THIRD EDITION REVIEWERS AND FOCUS GROUP PARTICIPANTS

Malikah Abdullah, Norfolk State University

Sherri Arthur Oklahoma City Community College

Mark Bland, University of Central Arkansas Glenn Cohen, Troy University

Gregory Dahlem, Northern Kentucky University

Michael (Thomas) Dodson, Mississippi University for Women

Dani Ducharme, Waubonssee Community College

Stefanie Gazda, University of Massachusetts, Boston

Jim Harper, Sam Houston State University

Tina Hopper, Missouri State University

Carl Johansson

Fresno City College

Wendy Kuntz, Kapi’Olani Community College

Brenda Leady, University of Toledo

Terry Miller, Central Carolina Community College

Sue Mungre, Northeastern Illinois University

Nikki Plaster, Golden West College

Bill Rogers, Ball State University

SECOND EDITION REVIEWERS AND FOCUS GROUP PARTICIPANTS

Shazia Ahmed, West Virginia University

Andrea Bixler, Clarke College

Terrence Boyle, Mississippi State University

Wendy J. Brown, Danville Area Community College

Sibyl Bucheli, Sam Houston State University

Jamie Burchill, Troy University

William Caire, University of Central Oklahoma

James Castle, Imperial Valley College

Yijing Chen, Kent State University

Tami Dahl, North Dakota State University

Greg Dahlem, Northern Kentucky University

Hattie Dambroski, Normandale Community College

Lewis Deaton, University of Lousiana H. Alan DeRamus, University of Louisiana

Danielle Ducharme, Waubonsee Community College

Denise Due-Goodwin, Vanderbilt University

Robert E. Farrell, Jr., Pennsylvania State University

Brandon Lee Foster, Wake Technical Community College

Samantha Furr-Rodgers, Stanly Community College

Kathy Gallucci, Elon University

Chunlei Gao, Middlesex Community College

J. Yvette Gardner, Clayton State University

Jenny Gernhart, Iowa Central Community College

Heather Giebink, Pennsylvania State University

Mary Gobbett, University of Indianapolis

Larry Gomoll, Stone Child College

Cara Gormally, Gallaudet University

Melissa Greene, Northwest Mississippi Community College

Melissa Gutierrez, University of Southern Mississippi

James Harper, Sam Houston State University

Mary Haskins, Rockhurst University

Jody Hawkins, College of Southern Idaho

Tina Hopper, Missouri State University

Joseph Daniel Husband, Florida State College, Jacksonville

David Rudge

Western Michigan University

Justin Shaffer University of California, Irvine

Debbie Sipes, Polk State College

Jennifer Stueckle West Virginia University

Andrew Swanson State College of Florida

Manjushri Kishore, Heartland Community College

Jennifer Kneafsey, Tulsa Community College

Wendy Kuntz, Kapi’Olani Community College

Brenda Leady, University of Toledo

Maureen Leupold, Genesee Community College

Jeffrey Mahr, Georgia Perimeter College

Lisa Maranto, Prince George’s Community College

Heidi Marcum, Baylor University

Bonique Morton, Youngstown State University

Vamsi Nalam, Indiana University–Purdue

University Indianapolis

James Newcomb, New England College

Lori Nicholas, New York University

ERIC J. SIMON, PH.D. New England College, Henniker, NH

Angel Nickens, Northwest Mississippi Community College

Mary O’Sullivan, Elgin Community College

Cassia Oliveira, Lyon College

Katherine Phillips, North Hennepin Community College

FIRST EDITION REVIEWERS

Shamili Ajgaonkar, Sandiford College of DuPage

Penny Amy, University of Nevada, Las Vegas

Kim Atwood, Cumberland University

David Ballard, Southwest Texas Junior College

Marilyn Banta, Texas State University

Patricia Barg, Pace University

David Belt, Metropolitan Community College, Penn Valley

Anna Bess Sorin, University of Memphis

Andrea Bixler, Clarke University

Susan Bornstein-Forst, Marian University

Randy Brewton, University of Tennessee, Knoxville

Peggy Brickman, University of Georgia

Steven Brumbaugh, Green River Community College

Stephanie Burdett, Brigham Young University

Greg Dahlem, Northern Kentucky University

Mary Dettman, Seminole State College of Florida

Vanessa Quinn, Purdue University

Eileen Roark, Central Connecticut State University and Manchester Community College

Peggy Rolfsen, Cincinnati State College

Carlos Santamaria, Sam Houston State University

Mark Schoenbeck, University of Nebraska

Justin Shaffer, North Carolina A&T State University

Clint Springer, St. Joseph’s University

Andrew Swanson, University of Arkansas

Eden L. Effert, Eastern Illinois University

Jose Egremy, Northwest Vista College

Hilary Engebretson, Whatcom Community College

Brian Forster, St. Joseph’s University

Brandon Foster, Wake Technical Community College

Thomas Gehring, Central Michigan University

Larry Gomoll, Stone Child College

Tammy Goulet, University of Mississippi

Eileen Gregory, Rollins College

David Grise, Texas A&M University–Corpus

Christi

Tom Hinckley, Landmark College

Kelly Hogan, University of North Carolina at Chapel Hill

Christopher Jones, Moravian College

Jacob Krans, Central Connecticut State University

Pramod Kumar, Northwest Vista College

Wendy Kuntz, University of Hawai’i

Dana Kurpius, Elgin Community College

Brenda Leady, University of Toledo

Maureen Leupold, Genesee Community College

Mark Manteuffel, St. Louis Community College

Debra McLaughlin, University of Maryland, College Park

Heather Miller, Front Range Community College and Kaplan University

Lisa Misquitta, Quinebaug Valley Community College

Pamela Monaco, Molloy College

Ulrike Muller, California State University, Fresno

Lori Nicholas, New York University

Monica Parker, Florida State College, Jacksonville

Don Plantz, Mohave Community College

Gregory Podgorski, Utah State University

Robyn A. Puffenbarger, Bridgewater College

Kayla Rihani, Northeastern Illinois University

FIRST EDITION CLASS TESTERS AND INTERVIEW PARTIPANTS

Leo Alves, Manhattan College

Tonya Bates, University of North Carolina at Charlotte

Brian Baumgartner, Trinity Valley Community College

Lisa Blumke, Georgia Highlands College

TJ Boyle, Blinn College

Michelle Brewer, Central Carolina Technical College

Melissa Caspary, Georgia Gwinnett College

Krista Clark, University of Cincinnati, Clermont

Merry Clark, Georgia Highlands College

Reggie Cobb, Nash Community College

Angela Costanzo, Hawai’i Pacific University, Loa Evelyn Cox, University of Hawai’i, West Oahu

Hattie Dambrowski, Normandale Community College

Kelsey Deus, Caspar College

Lisa Delissio, Salem State University

Suzanne Wakim, Butte Community College

Jennifer Wiatrowski, Pasco-Hernando College

Leslie Winemiller, Texas A&M University

Nancy Risner, Ivy Tech Community College

Bill Rogers, Ball State University

David Rohrbach, Northwest Vista College

Chris Romero, Front Range Community College, Larimer Campus

Checo Rorie, North Carolina Agricultural and Technical State University

Amanda Rosenzweig, Delgado Community College

Kim Sadler, Middle Tennessee State University

Steve Schwartz, Bridgewater State University

Tara Scully, George Washington University

Cara Shillington, Eastern Michigan University

Stephen Sumithran, Eastern Kentucky University

Suzanne Wakim, Butte Community College

Frances Weaver, Widener University

Susan Whitehead, Becker College

Jennifer Wiatrowski, Pasco-Hernando Community College

Matthew Wund, The College of New Jersey

Dani Ducharme, Waubonsee Community College

Jennifer Ellie, Wichita State University

Sachie Etherington, University of Hawai’i, Manoa

Christy Fleishacker, University of Mary

Brandon Foster, Wake Technical Community College

Valerie Franck, Hawai’i Pacific University

Jennifer Fritz, University of Texas, Austin

Kathy Galluci, Elon University

Chunlei Gao, Middlesex Community College

Mary Gobbett, University of Indianapolis

Erin Goergen, St. Petersburg College, Clearwater

Marla Gomez, Nicholls State University

Larry Gomoll, Stone Child College

David Grise, Texas A&M University, Corpus Christi

Melissa Gutierrez, University of Southern Mississippi

Barbara Hass Jacobus, Indiana University–Purdue University, Columbus

Debra Hautau, Alpena Community College

Jon Hoekstra, Heartland Community College

Tina Hopper, Missouri State University

Joseph Husband, Florida State College, Jacksonville

John Jenkin, Blinn College

Jamie Jensen, Brigham Young University

Julie Johns, Cincinnati State Community College

Anta’Sha Jones, Albany State University

Ambrose (Trey) Kidd, University of Missouri, St. Louis Manju Kishore, Heartland Community College

Cindy Klevickis, James Madison University

Tatyana Kliorina, Trinity University

Karen Koster, University of South Dakota

Barbara Kuehner, University of Hawai’i, West Hawai’i

Dana Kurpius, Elgin Community College

Jennifer Landin, North Carolina State University

Grace Lasker, Lake Washington Institute of Technology

Brenda Leady, University of Toledo

Sharon Lee-Bond, Northampton Community College

Ernest May, Kansas City Kansas Community College

MaryAnn Menvielle, California State University, Fullerton

Kim Metera, Wake Technical Community College

FIRST EDITION FOCUS GROUP PARTICIPANTS

Christine Andrews, Lane Community College

Morvarid Bejnood, Pellissippi State Community College

Cindy Bida, Henry Ford Community College

Nickolas Butkevich, Schoolcraft College

Susan Finazzo, Georgia Perimeter College

Jennifer Gernhart, Iowa Central Community College

Kristy Halverson, University of Southern Mississippi

Jody Hawkins, College of Southern Idaho

Wendy Jamison, Chadron State College

Kevin B. Jones

Charleston Southern University

Jacqueline Jordan, Clayton State University

Katrina Kirsch, Western Wyoming Community College

Heather Miller, Front Range Community College and Kaplan University

Pamela Monaco, Molloy College

Punya Nachappa, Indiana University–Purdue University, Fort Wayne

Kathryn Nette, Cuyamaca College

Betsy Ott, Tyler Junior College

Mary O’Sullivan Elgin Community College

Dianne Purves, Crafton Hills College

Peggy Rolfsen, Cincinnati State Community College

Checo Rorie, North Carolina A&T State University

Brian Sailer, Central New Mexico Community College

Daita Serghi, University of Hawai’i, Manoa Vishal Shah, Dowling College

David Smith, Lock Haven University

Patti Smith, Valencia Community College

Adrienne Smyth, Worcester State University

Wendy Stankovich, University of Wisconsin, Platteville

Jennifer Kneafsey, Tulsa Community College

Margaret Major, Georgia Perimeter College

Lisa Maranto, Prince George’s Community College

Cassandra Moore-Crawford, Prince George’s Community College

Ruben Murcia, Rose State College

Kim Sadler, Middle Tennessee State University

Frank Stanton, Leeward Community College

Olga Steinberg, Hostos Community College

Fengjie Sun, Georgia Gwinnett College

Ed Tall, Seton Hall University

Lavon Tonga, Longview Community College

Maria Trone, Valencia College, Osceola

Dan Trubovitz, San Diego Miramar College

Encarni Trueba, Community College of Baltimore County

Larchinee Turner, Central Carolina Technical College

Marty Vaughan, Indiana University–Purdue University, Indianapolis

Justin Walguarnery, University of Hawai’i, Manoa

Jim Wallis, St. Petersburg College, Tarpon Springs

Rebekah Ward, Georgia Gwinnett College

Jamie Welling, South Suburban College

Clay White, Lone Star College

Leslie Winemiller, Texas A&M University

Denise Shipley, Mountain View College

Kathy Watkins, Central Piedmont Community College

Christina Weir, Eastern New Mexico University, Roswell

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1 An Introduction to the Science of Life

The Chemistry of

3 The Cell: The Fundamental Unit of Life

3.1

3.3 Membranes are made from two layers of lipids

3.4 Membranes regulate the passage of materials

3.5 The nucleus houses DNA packaged as chromosomes

3.6 Several organelles participate in the production of proteins

3.7 Chloroplasts and mitochondria provide energy to the cell

3.8 Various cellular organelles provide storage, movement, and shape

Energy and Life

4.4 Photosynthesis occurs in two linked stages

4.5 In the light reactions, the energy of sunlight is captured as chemical energy

4.6 In the Calvin cycle, high-energy molecules are used to make sugar

4.7 In

4.8 Cellular respiration is divided into three

In

5 Chromosomes and Inheritance

5.1 Cell division provides for reproduction, growth, and repair 104

5.2 Chromosomes are associations of DNA and protein 106

5.3 Cells have regular cycles of growth and division 108

5.4 During mitosis, the nucleus of the cell divides 110

5.5 During cytokinesis, the cell is split in two 112

5.6 Nuclear transfer can be used to produce clones 114

5.7 Gametes have half as many chromosomes as body cells 116

5.8 Meiosis produces gametes 118

5.9 Mitosis and meiosis have important similarities and differences 120

5.10 Several processes produce genetic variation among sexually reproducing organisms 122

5.11 Mistakes during meiosis can produce gametes with abnormal numbers of chromosomes 124

5.12 Mendel deduced the basic principles of genetics by breeding pea plants 126

5.13 A Punnett square can be used to predict the results of a genetic cross 128

5.14 Mendel’s law of independent assortment accounts for the inheritance of multiple traits 130

5.15 Pedigrees can be used to trace traits in human families 132

5.16 The inheritance of many traits is more complex than Mendel’s laws

5.17 Linked genes may not obey the law of independent assortment

5.18 Sex-linked genes display unusual inheritance patterns 138

6 DNA: The Molecule of Life

6.1 DNA is a polymer of nucleotides 140

6.2 During DNA replication, a cell duplicates its chromosomes 142

6.3 DNA directs the production of proteins via RNA 144

6.4 Genetic information flows from DNA to RNA to protein 146

6.5 Transcription creates a molecule of RNA from a molecule of DNA 148

6.6 Translation involves the coordination of three kinds of RNA 150

6.7 Translation creates a molecule of protein via the genetic code 152

6.8 Gene expression is regulated in several ways 154

6.9 Signal transduction pathways can control gene expression 156

6.10 Mutations can have a wide range of effects 158

6.11 Loss of gene expression control can result in cancer 160

6.12 Cancer is caused by out-of-control cell growth 162

6.13 Genetic engineering involves manipulating DNA for practical purposes 164

6.14 DNA may be manipulated many ways within the laboratory 166

6.15 Plants and animals can be genetically modified 168

6.16 PCR can be used to multiply samples of DNA 170

6.17 DNA profiles are based on STR analysis 172

6.18 Whole genomes can be sequenced 174

6.19 Gene therapy aims to cure genetic diseases 176

7 Darwinian Evolution

7.1 Darwin’s influences and experiences led him to publish his theory of evolution 178

7.2 Unequal reproductive success leads to natural selection 180

7.3 Evolution affects our daily lives 182

7.4 The fossil record provides important evidence for evolution 184

7.5 Evidence for evolution is found in the natural world 186

7.6 Populations are the units of evolution 188

7.7 Evolution proceeds through several mechanisms 190

7.8 The geologic record ties together the history of Earth and its life 192

7.9 Macroevolution encompasses large-scale changes 194

7.10 Species are maintained by reproductive barriers 196

7.11 Speciation can occur through various mechanisms 198

7.12 Taxonomy is the classification of life 200

7.13 Phylogenetic trees represent hypotheses about evolutionary history 202

8 Biodiversity 1: Microscopic Organisms

8.1 Biologists hypothesize that life originated in a series of stages 204

8.2 Prokaryotes have unique cellular structures 206

8.3 Archaea are found in extreme habitats 208

8.4 Bacteria are very numerous and common 210

8.5 Bacteria can transfer DNA 212

8.6 Eukaryotic cells evolved from prokaryotic cells 214

8.7 Protists are very diverse 216

8.8 The origin of multicellular life was a major milestone in evolution 218

8.9 Viruses are nonliving parasites 220

8.10 HIV cripples the human immune system 222

8.11 Prions and viroids are nonliving parasites even smaller than viruses 224

9 Biodiversity 2: Fungi and Plants

9.1 Fungi are a diverse group of eukaryotes 226

9.2 Fungi have specialized structures and means of reproduction 228

9.3 Plants have unique adaptations that allow them to survive on land 230

9.4 Plant bodies consist of roots, stems, and leaves 232

9.5 Plant bodies follow a structural hierarchy 234

9.6 Four major groups of plants have evolved 236

9.7 Bryophytes are seedless, nonvascular plants 238

9.8 Vascular tissue transports water and nutrients 240

9.9 Ferns are seedless vascular plants 242

9.10 The first plants to evolve seeds were gymnosperms 244

9.11 Angiosperms dominate the modern landscape 246

9.12 Flowers, fruit, and seeds aid angiosperm reproduction 248

9.13 Angiosperms grow in length and in thickness 250

10

Biodiversity 3: Animals

10.1 Animals are multicellular consumers that evolved from colonial protists 252

10.2 Sponges and cnidarians have unusual body features 254

10.3 Three phyla of worms have unique structures and belong to separate evolutionary lineages 256

10.4 Mollusks are a very diverse phylum 258

10.5 The arthropods are extremely diverse and numerous 260

10.6 Echinoderms are a sister phylum to the chordates 262

10.7 Vertebrates belong to the chordate phylum 264

10.8 The first vertebrates to evolve were fishes 266

10.9 Amphibians and reptiles were the first tetrapods to occupy land 268

10.10 Mammals have hair and produce milk 270

10.11 Humans evolved from ancestral primates several million years ago 272

11 Human Body Systems

11.1 Animal bodies are organized into a structural hierarchy 274

11.2 The human body contains several major types of tissues 276

11.3 An animal’s internal environment remains relatively constant 278

11.4 The human digestive system consists of an alimentary canal and accessory organs 280

11.5 Food is processed in a series of stages 282

11.6 Proper nutrition provides energy and building materials 284

11.7 An unbalanced diet or malfunctioning digestive system can lead to health problems 286

11.8 The respiratory system exchanges gases between the environment and the body 288

11.9 The circulatory system transports materials throughout the body 290

11.10 The heart is the hub of the human circulatory system 292

11.11 Blood contains cells in liquid 294

12 Ecology

12.1 Ecology affects your life and our broader society 320

12.2 Ecology is the scientific study of organisms in their environments 322

12.3 Ecosystems include a variety of abiotic factors 324

12.4 Populations vary in age structure, survivorship, density, and dispersion 326

12.5 Growth models can predict changes in population size 328

12.6 Human population growth has been exponential 330 12.7 Interactions between species play important roles in communities 332

12.8 Food webs describe multiple trophic structures 334

12.9 Several factors affect species diversity 336

12.10 Invasive species can disrupt ecosystems 338

12.11 Biodiversity is measured on many levels 340

12.12 There are a variety of terrestrial biomes 342

12.13 Aquatic biomes cover most of Earth’s surface 344

12.14 Energy is lost as it moves through trophic levels 346

12.15 Elements cycle through the biosphere

12.16 All water on Earth is interconnected in a global cycle

12.17 People cause many ecological problems 352

11.12 The immune system contains a huge number of defensive elements 296

11.13 Immune system malfunctions cause a variety of disorders 298

11.14 The endocrine system regulates the body via hormones 300

11.15 The urinary system regulates water and rids the body of wastes 302

11.16 Males and females produce, store, and deliver gametes 304

11.17 A human develops from a single cell 306

11.18 Issues of reproductive health affect us all 308 11.19 The brain is the hub of the human nervous system 310 11.20 The nervous system receives input, processes it, and sends output 312 11.21 The senses use receptors to convey information about the outside world

11.22 The human skeleton contains 206 bones

Skeletal muscles produce movement

12.18 People can solve ecological problems

12.19 Human activity is causing a rise in greenhouse gases

APPENDICES

A: Metric Conversion Table A-1

B: The Periodic Table B-1

GLOSSARY G-1 CREDITS C-1 INDEX I-1

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great power of attorney from American citizens to their only American government—gives no power whatever to the states or their governments. Nevertheless, with amusement and absolute incredulity, we shall hear every statesman of 1917 and every lawyer of 1920 assume and act and argue upon the extraordinary concept that the Fifth Article of that Constitution is a great power of attorney to the state legislatures as attorneys in fact for the American citizens.

“We all know the severe scrutiny to which the Constitution was exposed—some from their own knowledge, others from different sources. We know with what jealousy, with what watchfulness, with what scrupulous care its minutest provisions were examined, discussed, resisted, and supported by those who opposed and those who advocated its ratification.” (4 Ell. Deb. 486.) So spoke Martin Van Buren in the Senate on April 7, 1826. We sit in the conventions which made the Constitution of which he spoke. We listen to every word that is said therein. We hear the Fifth Article explained by its worder, Madison, as nothing but a mode of procedure. From not one of the opponents of the Constitution, not even from Henry, do we hear a single word of attack upon the Fifth Article except as to the weakness of the mode of procedure which it provides for evoking the power of the “people” themselves, assembled in “conventions” of the very same kind, to withdraw from the one American legislature, the Congress, some enumerated power of the First Article which is found oppressive to individual liberty. On the contrary, we not only hear no single word of complaint that the Fifth Article or any Article gives one iota of power to the state legislatures, but the whole complaint of all the opponents of the Constitution which we do hear is that it practically destroys all existing ability and power of those state governments. Only a moment ago we have heard Henry ask: “If you adopt this Constitution, why continue the state legislatures at all?”

Anticipating the extraordinary concept which we are later to hear in 1917 and 1920, that the citizens of America by the Fifth Article made a collection of the state legislatures an omnipotent government over everything in America, including every individual right, we wonder if the constitutional thinkers of 1917 and 1920 remember that we are

sitting with a people who, just five years before the conventions in which they and we sit, ended an eight-year war to make certain that there never again should be a government of that kind in America, to make certain that they themselves should never be the “subjects” of any government or the citizens of any nation whose government should have even one power to interfere with individual liberty, except power of that kind granted directly by its citizens themselves.

It was Maclaine in the North Carolina convention who first used the exact expression that the Americans, who sit in the conventions where we are, were a people “better acquainted with the science of government than any other people in the world.” In the same convention, on July 29, 1788, this is what he had to say about the consistent attack upon the Constitution, because it robbed the state legislative governments of so much of their previous power. “Mr. Chairman, that it will destroy the state sovereignty is a very popular argument.... Government is formed for the happiness and prosperity of the people at large. The powers given it are for their own good.... The powers to be given the general government are proposed to be withdrawn from the authority of the state governments, in order to protect and secure the union at large. This proposal is made to the people. No man will deny their authority to delegate powers and recall them, in all free countries.... It may be justly said that it [the Constitution which contains the Fifth Article] diminishes the power of the state legislatures, and the diminution is necessary to the safety and prosperity of the people.” (4 Ell. Deb. 180.) It certainly would have surprised Maclaine, as well as all the Americans in those conventions, to have heard any one of themselves stating that the same Constitution vested the state governments with an omnipotence they had never possessed, the very omnipotence denied to the British Parliament eleven years earlier.

In the Virginia convention we hear Madison, who drafted and suggested the Fifth Article at Philadelphia, speak of the important distinction between the makers of the federal Articles of 1781, only seven years made, when we sit in that Virginia convention, and the “convention” makers of the proposed Articles of the new national Constitution. If these “conventions” make it, he says, it will be a

government established, not through the intervention of the legislatures but by the people at large. Fie goes on to say “In this particular respect, the distinction between the existing and proposed governments is very material. The existing system has been derived from the dependent derivative authority of the legislatures of the states; whereas this is derived from the superior power of the people. If we look at the manner in which alterations are to be made,” now referring directly to the Fifth Article, “the same idea is, in some degree, attended to.” (3 Ell. Deb. 94.)

We feel that it will be quite difficult, when we come later to the constitutional thinkers of 1917 and 1920, for them to convince us that Madison meant his Fifth Article to alter “the dependent derivative authority of the legislatures of the state” and, whenever another government makes the suggestion, put that dependent authority above what he calls “the superior power of the people.”

And we feel that these “constitutional thinkers” will find it impossible to convince us when we recall Madison’s other words, directly referring to his Fifth Article and the existing power of the people, mentioned therein by the word “conventions.” These are the words to which we allude: “Were it [his Fifth Article] wholly national, the supreme and ultimate authority would reside in the majority of the people of the Union, and this authority would be competent at all times, like that of a majority of every national society, to alter or abolish its established government.” It is Madison himself who puts the one word “majority” in italics. He does so to call attention to the fact that his Article leaves “the supreme and ultimate authority” in the people (named as “conventions” in his Article) but not necessarily capable of exercise by the majority in any constitutional manner He goes on to explain this very fact by saying that when the mode of procedure prescribed in his Article is read, it is found that “in requiring more than a majority, and particularly in computing the proportion by states, not by citizens, it departs from the national and advances towards the federal character.” (Fed. No. 39.)

In New York we find Hamilton, who seconded the suggestion of Madison’s Fifth Article at Philadelphia, almost immediately after he had stated that there would be no danger in permitting Congress to

propose amendments since “the final decision in the case will rest with the people.” As we recall, Hamilton said this when the tentative Fifth Article mentioned no one but the people, “conventions,” as the maker of future Articles, because he and Madison and their associates at Philadelphia, whose minds had so far been concentrated upon the national First Article, had not yet grasped the fact, later stated by Hamilton to be his conviction, that all future changes would probably relate “to the organization of government and not to the mass of its powers.” We are, therefore, interested to find Hamilton, in New York, on Friday, December 14, 1787, pointing out that “It has not a little contributed to the infirmities of the existing federal system that it never had a ratification by the . Resting on no better foundation than the consent of the several legislatures, it has been exposed to frequent and intricate questions concerning the validity of its powers.... The possibility of a question of this nature proves the necessity of laying the foundations of our national government [the First Article grant of national powers] deeper than in the mere sanction of delegated authority [referring directly to the state legislatures which are mentioned in the Fifth Article]. The fabric of American empire ought to rest on the solid basis . The streams of national power ought to flow immediately from that pure, original fountain of all legitimate authority.” (Fed. No. 22.) The capitals are those of Hamilton himself. We rather feel that his stress upon the “” sharply contrasted with the state “legislatures,” as the only legitimate direct source of national power, such as is granted in the First Article and the Eighteenth Amendment, will be somewhat of a shock to the “constitutional thinkers” of 1917 and 1920. Sitting in the conventions of old, we rather recognize the capitalized words, where Hamilton says that national power in America “ought to rest on the solid basis of the ,” as a direct echo from the Statute of ’76, enacted only eleven years before those conventions. That Statute says that every just power of government must be derived directly from the governed.

And we become rather convinced that Hamilton and Madison, when submitting the Fifth Article at Philadelphia, never worded it so that national power in America could be granted through the

illegitimate authority of the state legislatures, when we read what either one or both of them have to say on Tuesday, February 5, 1788, in The Federalist, No. 49. “As the people are the only legitimate fountain of power, and it is from them that the constitutional charter, under which the several branches of government hold their power, is derived, it seems strictly consonant to the republican theory to recur to the same original authority”—a direct reference to the “conventions” of the Seventh and the Fifth Articles—“whenever it may be necessary to enlarge, diminish, or new-model the powers of government.”

If we ever had any doubt as to what Hamilton meant the Fifth Article to provide, our doubt is ended forever when we hear Hamilton, in the New York Convention, speak of the state legislatures, which the “constitutional thinkers” of 1917 and 1920 assume to have been made attorney in fact for the American people for every purpose by that Fifth Article. “Look through their history,” he says, speaking of those state legislative governments. “What factions have arisen from the most trifling causes! What intrigues have been practiced for the most illiberal purposes! Is not the State of Rhode Island, at this moment, struggling under difficulties and distresses, for having been led blindly by the spirit of the multitude? What is her legislature but the picture of a mob?” Most of the states “are either governed by a single democratic assembly, or have a senate constituted entirely upon democratic principles. These have been more or less embroiled in factions, and have generally been the image and echo of the multitude.... Let us beware that we do not make the state legislatures a vehicle in which the evil humors may be conveyed into the national system.” (2 Ell. Deb. 317.) When Hamilton knew that these state legislatures were not the legitimate source of national powers in a republic and when he had this view of their general character, will any sane man believe that he advocated that the citizens of America should make these legislatures (although the citizens of America elect not a single member in them) absolute attorney in fact for the citizens of America for all purposes? Will any sane man believe that he proposed to substitute them, as grantors of national power, for the “conventions” of the Fifth Article, which “conventions” already excited the admiration of the entire world,

according to the author of the Fifth Article, as the only safe or effective mode in which the free citizens of a nation could vest its government with any power to interfere with their own individual liberty.

“The authority of Constitutions over governments, and the sovereignty of the people over Constitutions, are truths which are at all times necessary to be kept in mind; and at no time, perhaps, more necessary than at present.” This impressive statement of the truth that the “governments,” state “legislatures,” never were placed by the Fifth Article above the Constitution in which it is, and above the sovereignty of the people, mentioned therein as “conventions,” and cannot be placed there by another government, the Congress, is not a statement made in the heat of the controversy about the Eighteenth Amendment. It is the statement of James Madison, author of the Fifth Article, made in the Virginia House of Delegates in 1799;

It is becoming almost impossible for us, as we sit in these conventions and hear every word that is said, to understand the source of the modern thought, if we can dignify it by calling it “thought,” that the Fifth Article is a power of attorney from the citizens of America to the state governments, every member of which is elected by the citizens of the respective states and not by the citizens of America. It is beginning to grow upon us that any such “thought” is based on sheer assumption and that the entire record of the “conventions” is a closed book to those who hold the assumption. We know that they have the explicit statement of the Tenth Amendment, that the Constitution gives no power of any kind to any donee except the one new American government, the government with the enumerated powers of the First Article. As no power of attorney was ever written or can be conceived except one which grants the attorney in fact some power, the Tenth Amendment makes absolutely certain that neither the Fifth Article nor the entire Constitution gives to the state legislatures any power as attorneys in fact for the citizens of America.

Moreover, breathing the atmosphere of those first “conventions” of the kind named in the Seventh and the Fifth Articles, the

“conventions” where individual liberty of the American is the only object of advocate and opponent of the Articles under consideration, we begin to sense that the holders of the impossible assumption have never fully grasped the amazing and vital distinction between “state legislatures” and “conventions” of the kind named in the Seventh and the Fifth Articles. When we shall hear the “constitutional thinkers” of 1917 and 1920 speak of the “legislatures” and the “conventions” as two different agents given omnipotent attorneyship in fact over all the individual rights of the citizens of America, we shall wonder if these thinkers appreciate that the “state legislatures” are permanent bodies, always existing, and that the “conventions” of the Fifth Article are, to the “conventions” in which we sit, bodies that never would have an existence until some future moment, when the American citizens themselves would again be called to assemble in and thus make those “conventions.” Nothing could show more clearly that the “conventions” of the Seventh Article looked upon the “conventions” of the Fifth Article, not as the donee of any power of attorney, but as themselves or their posterity, the citizens of America, assembling again to determine whether there shall be any change in the distribution of power to interfere with their individual liberty. And our thought, re-echoed again and again by Marshall and others from the Bench of the Supreme Court in the century that follows the first assembling of these “conventions,” seems but the repetition of what we hear said in the Massachusetts Convention as the tribute of its Americans to the Fifth Article.

On January 23, 1788, the Americans, assembled in Massachusetts, took up the consideration of that Article. As in every convention, there had been great opposition to the earlier Articles; as in every convention, nearly all of it had been to the great national powers of the First Article granted to the new government and taken from the state governments; and, as in every convention, almost all of this opposition had been the continued complaint that the state governments were being destroyed.

It was not that the Americans loved the state governments. The truth is that, like every natural human being, they objected to all governments. Their sole thought was fear of oppressive government

infringement upon their individual liberty In this respect, the Americans in each convention feared their own legislative government less than the new proposed Congress, because they would elect all the members of the former and only a few members of the latter. If it had been suggested, by any of the many opponents of the new Constitution, that any possible twisting of the words of the Fifth Article meant that governments outside their state, not one of whose members they themselves would elect, could infringe upon their every individual right, without any constitutional restraint, the record of every convention would have been one unanimous “,” against the new Constitution. But, as no “constitutional thinker” of 1917 or 1920 sat in any of those conventions, no such suggestion was ever made therein.

And so, on that January 23, we hear the Fifth Article read in that Massachusetts convention, and we see Rufus King rising and we hear him state that “He believed gentlemen had not, in their objections to the Constitution, recollected that this Article was a part of it; for many of the arguments of gentlemen were founded on the idea of future amendments being impracticable.” He dwelt “on the superior excellence of the proposed Constitution in this particular, and called upon gentlemen to produce an instance, in any other national constitution, where the people had so fair an opportunity to correct any abuse which might take place in the future administration of the government under it.”

And then we hear Dr. Jarvis: “Mr. President, I cannot suffer the present Article to be passed, without rising to express my entire and perfect approbation of it. Whatever may have been my private opinion of any other part, or whatever faults or imperfections I have remarked, or fancied I have seen, in any other instance, here, sir, I have found complete satisfaction: this has been a resting place, on which I have reposed myself in the fullest security, whenever a doubt has occurred, in considering any other passage in the proposed Constitution. The honorable gentleman last speaking has called upon those persons who are opposed to our receiving the present system, to show another government, in which such a wise precaution has been taken to secure to the people the right of

making such alterations and amendments, in a peaceable way, as experience shall have proved to be necessary. Allow me to say, sir, as far as the narrow limits of my own information extend, I know of no such example. In other countries, sir,—unhappily for mankind,— the history of their respective revolutions has been written in blood; and it is in this only that any great or important change in our political situation has been effected, without public commotions. When we shall have adopted the Constitution before us, we shall have in this Article an adequate provision for all the purposes of political reformation. If, in the course of its operation, this government shall appear to be too severe, here are the means by which this severity may be assuaged and corrected.... We have united against the British; we have united in calling the late federal convention; and we may certainly unite again in such alterations as in reason shall appear to be important for the peace and happiness of America.” (2 Ell. Deb. 116.)

No man ever voiced such sentiments, no conventions of Americans ever listened to them, with any knowledge or thought that the Fifth Article, “the wise precaution” to secure the liberty of the individual if the government with the national powers of the First Article oppressed that liberty, was itself a grant to another government, ten legislatures outside of the Massachusetts in which that convention was held, to infringe upon the individual liberty of every American in Massachusetts on every subject without any constitutional restraint.

And so, we average Americans end our education in the only “conventions,” named in the Seventh or the Fifth Articles, which yet have assembled. And we end that education knowing that there is nothing anywhere in the Constitution those conventions adopted, and especially nothing in the Fifth Article, which changed the free American into a subject of any government or governments in America. Everything we have heard—and what we have repeated is but little of what we have heard—serves but to emphasize the only meaning of its “apt, precise and classic English,” the plain meaning which we got from its language when we read it at the beginning of these conventions with the Americans who made it.

It is, as its author explained it, naught but a constitutional mode of procedure in which may be thereafter exercised, in a constitutional manner, either the limited ability of state governments to make Articles which do not concern themselves with the infringement of individual liberty or the unlimited ability of the people themselves, the “conventions” of the kind in which we have sat, to make any Articles. The procedure prescribed for such constitutional exercise is simplicity itself to those who sit in those conventions. It is exactly the procedure just followed (up to the point where the work of any proposer of a new Article and its mode of ratification must end) by the Philadelphia Convention which drafted it and the other six Articles. The Philadelphia Convention found itself without any mode of procedure in which could be evoked to exercise the existing and exclusive power of the people of America to grant any government power to infringe upon the individual liberty of the American citizens. There being no constitutional mode of procedure, no designated body to draft Articles with such grants and to propose them and to ascertain and propose the valid mode of ratification for them, the Philadelphia Convention did that work, guided only by basic American doctrine, the Statute of ’76 and the experience of the “conventions” which had made the national Articles of 1776. It followed a certain mode of procedure in the doing of these things, knowing and stating that to draft Articles and propose them and ascertain and propose the right mode of ratification for them is not the exercise of any power. With a knowledge which we of a later generation never should have forgotten—and which we who have been educated with them never will forget—the Philadelphia Convention knew that there were two makers of Articles in America, each of which had exercised its respective and different ability to make them, during the eleven years which preceded the Philadelphia Convention. They knew that every Article that was national could be made by no one but the people themselves, the “conventions,” which had made the national Articles of ’76 and which are named as the makers of all future Articles of that kind in the Seventh and the Fifth Articles proposed by Philadelphia. And so, when the Philadelphia Convention had drafted its Articles and was about to propose them, it recognized the legal necessity of

ascertaining, from the nature of those Articles, whether they were in the power of both or only of one of those existing makers of Articles. In the ascertainment, with their minds on the First Article grants of national power to interfere with individual liberty, they knew that no governments in America could make an Article of that kind. Their ascertainment was then ended and they knew that they must propose that mode of ratification which would send their Articles to the only valid ratifiers, the people themselves, the “conventions” of the Seventh and the Fifth Articles.

This was the procedure they had followed, when there was no constitutional mode of procedure provided. And so, with the extraordinary wisdom that characterized everything they did, that Philadelphia Convention wrote exactly the same procedure into the Fifth Article so that never again there might be lacking in America a constitutional mode of procedure for the evoking and the exercise of the only power that is ever exercised when constitutional Articles are made, the power of making them. As the Philadelphia Convention ended its existence with its own proposals, some new body had to do that work, when any new Article was to be proposed. As the work of the Philadelphia Convention had not been the exercise of any power but merely the work of proposing, it was a certainty that the new constitutional mode, exactly the same mode as that of Philadelphia, would also be the exercise of no power. And so, the Philadelphia Convention named the Congress (or a convention demanded by the state legislatures) to do the work of the Philadelphia Convention in drafting and proposing any new Article, and it named the Congress to perform the duty of ascertaining (by the nature of the new drafted Article) which of the two makers could make it, and then to propose a mode of ratification by which it would be validly ratified by such competent maker. As to the only powers ever to be exercised in the making of any new Article, the power of legislatures to make federal Articles, and the exclusive power of the people or “conventions” to make national Articles, the constitutional mode of procedure did not (nor could it, if Americans were not to become “subjects”) give the governments any of the exclusive ability of the people or “conventions,” and it did not (nor could it, if America were to be a republic) alter the existing ability of the majority of the

American people to make their governments what they will. But, for the very practical purpose which Madison so clearly explained, the purpose of providing some check upon the tyranny of the majority or an aggressive minority over the individual rights of all Americans, the Fifth Article procedure could and did fail to provide any method in which government power to interfere with individual liberty, as all surrendered power of that kind was distributed between different governments in the Constitution, could be changed in any way or transferred from one government to another, unless the “conventions” of the American citizens in three fourths of the states said “Yes” to any proposed change or transfer.

The Philadelphia Convention having proposed this particular check upon the existing ability of the people themselves to oppress individual liberty, a check which makes the words “by conventions in three fourths thereof” by far the most important words, the Fifth Article goes on to prescribe exactly the same check on the exercise of the ability of the state legislatures to make federal Articles.

That the Fifth Article, a constitutional mode of procedure for the exercise of two different existing abilities, was not a grant of any power to the state legislative governments is something that was known to every man in the conventions which made that Fifth Article.

In the Pennsylvania convention, Wilson plainly stated the knowledge of all that the supreme power “resides in the people, as the fountain of government; that the people have not—that the people meant not—and that the people ought not—to part with it to any government whatsoever. In their hands it remains secure. They can delegate it in such proportions to such bodies, on such terms, and under such limitations, as they think proper. I agree with the members in opposition, that there cannot be two sovereign powers on the same subject.... My position is, sir, that, in this country, the supreme, absolute, and uncontrollable power resides in the people at large.” (2 Ell. Deb. 456 et seq.)

When more than half a century had passed, the same thing was known to those who knew American Constitutional Law.

“It is obviously impossible for the whole people to meet, prepare and discuss the proposed alterations, and there seems to be no feasible mode by which an expression of their will can be obtained, except by asking it upon the single point of assent or disapproval. But no body of representatives, unless specially clothed with power for that purpose by the people when choosing them, [ergo, no permanent state governments or legislatures] can rightfully take definitive action upon amendments or revisions; they must submit the result of their deliberations to the people— —for ratification or rejection.”

So spoke the great Cooley in reference to making changes in national constitutions in his work on Constitutional Limitations (7th ed., 1903, at p. 61).

When one hundred and seventeen years had passed since the conventions in which we just sat, the same thing was known in the Supreme Court, in 1907.

The powers the people have given to the General Government are named in the Constitution, and all not there named, either expressly or by implication, are reserved to the people and can be exercised only by them, or upon further grant from them. (Justice Brewer in Turner v. Williams, 194 U. S. 279.)

CHAPTER XIV

SEVENTEEN

ARTICLES RESPECT HUMAN FREEDOM

“For my own part, I acknowledge a thorough conviction that Amendments which may, upon mature consideration, be thought useful, will be applicable to the organization of the government, not to the mass of its powers.” (Fed. No. 85.)

This was the statement of Hamilton to the American people when he was asking them, about to assemble in their conventions, to make their First Article grants of enumerated powers to interfere with their individual freedom and to make their Fifth Article mode of procedure, in which they could exercise their exclusive ability, assembled again in “conventions,” to add or subtract from that grant of enumerated powers of that kind.

When he used the words, “mass of its powers,” he referred directly to that First Article grant. It contains all the powers that were ever given to any government to interfere with the individual freedom of the American citizen. He knew what the Supreme Court clearly declared in United States v. Cruikshank, 92 U. S. 542, that, beyond the scope of its enumerated powers, there is no government of the American citizens. He knew that, in the geographical territory which is each state, there would thereafter be two governments, the government of enumerated powers, governing the American citizens in that state under the First Article grants from the American citizens, and the state governments, governing the citizens of the state, under whatever grants of national power its state citizens gave that government.

The two governments in each state stand in their respective spheres of action in the same independent relation to each other, except in one particular, that they would if their authority

embraced distinct territories. That particular consists in the supremacy of the authority of the United States when any conflict arises between the two governments. (Justice Field, in Tarble’s case, in the Supreme Court, 13 Wall. 397.)

With such knowledge, he would have found it impossible to make the blunder of assuming that either government could give or join with governments in giving to the other any power to interfere with the citizens of the grantee in their enjoyment of individual liberty. He knew that, “In our republican forms of government, the absolute sovereignty of the nation is in the people of the nation; and the residuary sovereignty of each state, not granted to any of its public functionaries, is in the people of the state.” (Story, 1 Ell. Deb. 65.)

When Hamilton stated, in those days of 1788, his conviction that new Articles would relate to the “organization of government and not to the mass of its powers,” he was epitomizing the common knowledge of everyone, at that time, that the new Constitution was both federal and national. He was predicting that the power to interfere with individual liberty, to the extent which Americans had surrendered any power of that kind to their governments, had been so wisely distributed, between the government of the citizens of America and the political entity which was each state, that it would never be necessary for the citizens of America to alter that distribution by giving any more to their government. That is why he prophesied that no new Article of the kind which only “conventions” of those citizens can make, Articles changing the “mass of powers” granted to that government by the First Article, would ever be needed.

And we recall that this conviction probably influenced the last day remembrance in the Philadelphia Convention by Madison and himself, that the Fifth Article should also contain some reference to the state legislatures and their future exercise of their existing abilities to make federal or declaratory Articles, which do not change the “mass of powers” in that First Article grant and do not relate to government interference with the individual liberty of the American citizen.

The statement of Hamilton was not meant as a prophecy It was the result of a great mind reasoning from cause to probable effect. As we come down through the century or more that follows that statement and reach the beginning of the year 1917, we pay our tribute to a mind which could reason so correctly as to what would happen for more than a hundred years. We shall find that never again, from 1787 to 1917, did experience find it necessary that the “conventions” of the American citizens, the “conventions” named in the Seventh and the Fifth Articles, should again be assembled to alter, as they alone can alter, the mass of enumerated powers to interfere with their own individual liberty, which they granted in that First Article.

It is a striking commentary upon the ability of Hamilton and his generation, when contrasted with our modern “constitutional thinkers,” that he knew and appreciated what would happen in a century to come, while they have not understood what had happened in a century that had gone, although the record of that century was spread out before them to read. None of our leaders have appreciated the fact that every one of the first seventeen Amendments to our Constitution was of the kind that state legislatures had ability to make before our Constitution was even drafted at Philadelphia, because all were federal (or declaratory) Articles and were not national Articles. If they had known, after these seventeen Amendments were history, what he prophesied before the Constitution itself was adopted, the story of the last five years might not have been what we shall learn hereafter that it was.

That we may know, of our knowledge, what he prophesied and what they entirely overlooked and ignored, let us briefly examine the nature of those seventeen Amendments. Thus we will learn why governments, known at Philadelphia in 1787 to be incompetent to make national Articles, could and did make those seventeen Articles in the constitutional mode of procedure which the Fifth Article provided for the exercise of their existing ability limited to the making of federal or declaratory Articles.

The first ten Amendments were declarations insisted upon by the American citizens, assembled in “conventions” where we have sat,

as specific security against government usurpation of power over their individual freedom. The Supreme Court has repeatedly declared that everything in those ten Amendments was in the Constitution when it was ratified by those “conventions” and that the Amendments simply declared what the will of the people themselves, assembled in those “conventions,” had already established as the fundamental law in America. Certainly no man would challenge the prophecy of Hamilton on the theory that any one of those Amendments added to the mass of government powers to interfere with human freedom. They are, each and every one of them, the declaration that government cannot do “this” and government cannot do “that.” So far from being national Articles, the kind which only “conventions” of American citizens could or can make, the kind which tell government that it can command the American citizens on this or that subject, they are all Articles which tell government that it cannot command the American citizen. And, for the further security of the individual freedom of the American citizen, a security never needed more than in this year 1923, the Tenth Declaration emphatically declares who it is that the exclusive ability to alter, in one iota, “the mass of powers” (over the individual freedom of the American citizen) granted by them in the First Article and the exclusive ability to exercise any power of that kind over them, which was not granted in the First Article. Who can deny that, stated in our own words, this is what we find in the plain declaration of the Tenth Amendment? “We, the people of America, assembled in our conventions, have granted to the American government enumerated powers of the First Article. They are the only powers of that kind delegated to any government, by which it can interfere with our individual freedom in our capacity as American citizens. All powers, which the citizens of each state have hitherto had and which we have not taken from them herein, we have left with them; and the citizens of each state can grant so much of said powers as they please to their own government to govern them as citizens of that state. All other powers, outside those we have granted to our government to interfere with us and those we have left to the citizens of each state for their own respective exercise, we reserve exclusively to ourselves, in our capacity as citizens of America. And,

if any government should deem it wise that any one of these powers (which we so reserve exclusively to ourselves) should be exercised, we have provided in the Fifth Article the mode of procedure in which we, assembled in our conventions, can exercise it or grant it to the government which wants to exercise it.”

The powers the people have given to the General Government are named in the Constitution, and all not there named, either expressly or by implication, are reserved to the people and can be exercised only by them, or upon further grant from them. (Justice Brewer in Turner v. Williams, 194 U. S. 279.)

Later herein we Americans, fresh from the “conventions” which demanded these declarations for the better protection of individual liberty, are going to hear expounded a doctrine, which would mean that these declarations were themselves the American Magna Charta. We are going to hear, to our amazement, that these declarations were a compact between an omnipotent government and its subjects, ourselves, who have always imagined that we were “citizens” and not “subjects.” We are going to hear, to our utter incredulity, that these declarations are the promise of a government —itself omnipotent over ourselves and our Constitution and our government which we thought to be the supreme government in America—that we, the subjects of that omnipotent government, may have certain privileges which this omnipotent government will not take from us.

When first we shall hear this 1917 resurrection of the Tory concept that government is master and Americans are “subjects,” that government is the state and we are its assets, we shall naturally be astounded and indignant that even a few inhabitants should still retain what Madison called “the impious doctrine” of the Old World, that people are made for governments, not governments for the people. Quickly, however, we shall become alarmed to find how wide-spread, among the “constitutional thinkers” of our own generation, is the complacent acceptance of the Tory concept and to learn that, when its application in any given instance injures a client of one of those constitutional thinkers, no matter how he may argue

against the application in the particular instance, he neither knows nor uses for his client the fact that the Statute of ’76 repealed the doctrine on which Magna Charta rested, the Tory concept that government is the state and that we are its “subjects.”

It will be difficult for us to understand these leaders of our generation. In addition to what we have just lived through with the early Americans, these leaders will have before them countless decisions of the Supreme Court, flatly contradicting the Tory concept on which all these leaders reason. “The concessions of Magna Charta were wrung from the King as guarantees against the oppressions and usurpations of his prerogatives.... The omnipotence of Parliament over the common law was absolute, even against common right and reason. The actual and practical security for English liberty against legislative tyranny was the power of a free public opinion represented by the Commons.” We are going to find many of our leaders with the fixed Tory concept that there is in America a legislative government, unknown to the generation of Americans who created all government in America, and that this omnipotent government can do whatever it pleases, without any constitutional restraint, with almost everything in America, including ourselves and our Constitution and our American government. Curiously enough, we are going to be told that the one thing, in which this omnipotent legislative government falls short of absolute omnipotence, has nothing to do with any individual human freedom, but is its inability to change the equal representation of every state in our Senate. It will be quite amazing to us to hear these men, not proving this Tory concept but stating it as axiomatic American law and on it basing their every argument and audaciously claiming that our Supreme Court has no right even to consider whether it is so or not. But there will be something to alarm us even more. We will find our other “constitutional thinkers,” by circumstances obliged to challenge some particular deduction from this Tory concept, without any knowledge of the invincible facts which challenge the concept itself. To most of these men it will be as if the Statute of ’76 had been repealed and the result of our Revolution reversed, in 1787, by the very “conventions” in which we have sat. It will be our amazement to hear that these “conventions” never created the citizen of America

but that the American people, assembled therein, voluntarily made themselves “subjects” of the omnipotent government. And these modern “constitutional thinkers” will discuss this omnipotent government on the basis that those “conventions” made that particular government (not the one to whom the First Article grants its powers) attorney in fact for the general people of America, authorized to interfere with their individual freedom, “on all matters whatsoever.” To most of these men, it will be as if the Supreme Court, in countless ways, had not declared:

In this country, written constitutions were deemed essential to protect the rights and liberties of the people against the encroachment of power delegated to their governments, and the provisions of Magna Charta were incorporated into bills of rights.... Applied in England only as guards against executive usurpation and tyranny, here they have become bulwarks against arbitrary legislation; but in that application, as it would be incongruous to measure and restrict them by the ancient customary English Law, they must be held to guarantee, not particular forms of procedure, but the very substance of individual rights to life, liberty and property.

Of course, fresh from the conventions of old, we shall know that these men are talking nonsense. We shall know that the last two quoted statements from the Supreme Court, in Hurtado v. California, 110 U. S. 516, are but the expression of what was once the common knowledge of a people then “better acquainted with the science of government than any other people in the world.” But, as our knowledge shall be as the knowledge of the Americans of old, who fought an eight year war of sacrifice to make any such Tory concept in America absolute nonsense, it is well for us, while we are examining the nature of the first seventeen Amendments of our Constitution, to keep in mind one thing, which we now know and which, if it had not been forgotten, would have prevented the monumental blunder of the generation in which we ourselves are the American citizens. This one thing is the fact that the Constitution, made in the conventions we have just left, is a federal and a national Constitution. That fact and its influence upon the Convention, which

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