'I i
I
II f/Je,aop•fYK M. 11. *, HaropHIOK T. B., Tapac/OK c. 11., rna3KO B. 11. *XepCOHCK~~ rOCYABPCTB8HH~~ arpapHW~
YHMB8PCMT8T, r. XepcoH, yn. P03W noKCeM6ypr, 23, 73006, ~HCT~TYT arpo3Konor~M M6~oTexHonor~~ YAAH, YKpa~Ha, r. K~es, yn. MeTponor~YecKaR, 12, 03143
YKpa~Ha,
non~MOP$~3M
rEHET~K0-6~0X~M~4ECK~X C~CTEM
Y 3X~HAUE~ nYPnYPHO~ ~ 3X~HAUE~ Y3KOR~CTHO~ BmKHOCTh KOHTpOJUI reHeTWiecKoi1 crpyKTyphl pa3JI.l1IIHbiX BM.n,os neKapCTBeHHhiX pacTeHim .D,OCTBTO"'IHO O"'ICB.l1.D,HB, fJOCKOJibKY OHM
~BJU!IOTC~
HCTO"'l-
HHKOM CCTCCTBCHHbiX KOMITOHCHTOB (.n,eHCTBYIO~HX Haqan) MHOfHX Jieqe6HbiX npenapaTOB. oe3 rny6oKoro H3yqeHH~ CTPYKTYPhl .11 6HOXHMH'ICCK.l1X rrpoQecCOB,
npOHCXO)l~~HX
B
paCTCHH~X
HCB03MO)J(H0 ITOJIHOCTbiO npOX\YKTHBHO HC-
fJOJib30BBTb see HX QeHHbiC CBOHCTBa. 0}:(H.l1M .113 MeTO}:(OB OUCHKH oco6eHHOCTCH Hx BHJlOBOH H BHyrpHB.l1.D,OBofi reHeTMqecKoi:f CTPYKTYPhi ~BJUieTC~ H3Y'1CHHe fJOJTHMOp¢H3Ma fCHeTHK0-6.110XHMHqecKHX CHCTCM, Henocpe.D,CTBCI!HO
cB~3aHHhiX
c
¢YHX\aMeHTaJihHhiMH
BHYTPHKJieToqHhiMH
6HOXHMHqecKHMH
rrpoueccaMH. Mo)J(HO O)J(H}:(BTh, "'ITO BhffiBJienue YXI06HhiX noJIHMop¢HhiX reneTHKo-6uoxHMHqecKHX CHCTCM ll03BOJI~T KOHTpOJIHpOBBTh }:(HHBMJ1qecKHe IlpOUCCChi, npOHCXOJUI~He B llOllYJI~QH~X JJCKBpCTBCHHbiX pacTCHHH B pa3HbiX yc-
JTOBH~X pa1Be}:(eHH~, MapKHposaTh rpynnhr pacTeHHH BHYTPH nonyn~QHH, oTJmqaJO~uec~
fJOBhillJCHHhiMH npOX\YKTHBHhiMH xapaKTCpHCTHKBMH.
Oco6yJO axTyaJihHOCTh 3TO HMeeT .11 X\JI~ npe}:(cTaBHTenefi po}:(a 3XHHaQe~ (Echinacea Moench), oco6eHHO TaKHX, KaK 3XHHaQe~ 6ne}:(H~ H 3XHHaQe~ y3KOJIHCTHal!. 3TH BH}:(hl qaCTO nyTaiOT, KBK IIpH fJOCTBBKe HX CCMCHHOfO MBTCpHaJia, TBK .11 llpH npO}:(B)J(C Cblpb~
(5).
,lVJll TOTO, IIT06bi HMCTb B03MO)J(HOCTb BCCTH fJOHCK TBKHX CHCTeM, HB nepBOM 3TBlle He06XO}:(HMO HCCJie}:(OBBTb BH}:(OBbie, TKBHecneQH¢HqHhie OC06eHHOCTH 3KCllpCCCHH aJIJieJihHbiX BBp.11BHTOB reHeTHK0-6HOXHMJ1qeCKHX CHCTCM, fJOCKOJihKY B 3TOM OTHOIIICHHH B.l1.D,bl pO)lB
3XHHBUe~ OCTBJOTC~ )J,O
CHX
flOp HC)J,OCTBTOLfHO HCCJie)J,OBBHHb!MH.
l1
CCJIH B fJOCJie}:(Hee BpeM~ TBKHe pa60Tbl HBLfHHBIOT pa3BeJJfbiBBTbC~ B
ClllA (4),
TO )J,JJ~ CTpaH
CH,U
I1 oaJITHH HaM H3BeCTHa JIHIII OL(Ha ny6JIHKai~H~
KBCBJOIIIB~C~ BbffiBJICHH~ IlpH fJOMO~H .1130¢epMeHTOB fCHCTHqecKOll li3MeHql1BOCTH HBH6onee pacnpOCTpaHeHHOfO B KYJII>Type BH)J,B- 3XHHBUCH nypnypHOll
103
Ex libris ТРАВЫ ЖИЗНИ: http://medherbarium.blogspot.com
(3). Y KOTOpOH 6b!JIO HJyY.JeHO 14 cpepMeHTaJihHhiX CHCTeM, JIH!Ub B J(BYX 113 HHX MOJieKymrpHM MHO)!(eCTBeHHOCTb He 6biJia 06Hapy)!(eHa. Heo6xonHMO nonqepKHYTh, 'ITO K uacTOHIUeMy speMeHH B M0,1eKynHpHOH reHeTI1Ke pmpa6oTaHO MHOro MeTO]lOB, ll03BOJIHIOIUI1X MapKHpOBaTb 113MCH'li1BOCTb reHCTI1Y.JeCKOfO MaTep11aiia. 0J(HaKO LlO CI1X !lOp HaH60Jiee yno6Hb!M MeTOLI,OM Ll,JIH O)lHOBpeMeHHOH OQeHKH llOJIHMOpcpY13Ma fOMOJIOrH'lHbiX y pa3HbiX Bllll,OB CTpyKTypHbiX reHOB, CTporo COOTBCTCTBYIOJ.UHX fl0J111MOpcp113MY OT)lCJlbHbiX cpeHOB C OTHOCI1TeJibHO H3YY.JeHHOH 6HOXHMI1'leCKOH cpyHKQ11eH, OCTaeTCH MeTOLI, aHaJIHJa reHCTHY.JeCKH LJ,CTepMHH11pOBaHHOfO llOJIHMOpcp113Ma
3TOH CBHJH B HacTOHIQeii pa6oTe, c uen.hiO l13yY.JeHHH BHYTPHBH.UOBhiX H
Me)!(BllLI,OBhiX
OC06CHHOCTeH
fCHCTWICCKH
LI,CTCpMHHHpOBaHHOfO
llOJIHMOp-
cpH3Ma y JieKapCTBCHHhiX paCTCHHH, BhlllOJTHCH cpaBHHTCJlhHhiH aHaJII13 I10JillMOpcpH3Ma cpepMeHTOB o6mero BHYTPHKJTCTOY.JHoro MeTa6onHJMa B Kopr-mx, JIHCThHX, CTC6JUIX H DJIOJ(ax J(Byx BHJ(OB 3XHHaQCH purpurea
(L.) Moench) 11 YJKOJIHCTHOH (E.
nypnypHOH (Echinacea
3KcnepHMeHTaJlhHhiH MaTepHan .llJ1H J(aHHOH pa60Tbl llOJIYY.JCH OT pa3HbiX Ka'leCTBe MOJieKynHpHO-fCHCTH'leCKHX MapKepOB .llJ1H ODHCaHHH riOJil1-
MOpcp113Ma paJJIH'lHhiX BU.UOB 3XI1HaQeH Mhl paccMarpHBaJIH JIOKYChl, KO,LIHpYIQIQHe pH.Ll cpepMeHTOB
(7
reneTHKo-6MOXHMH'lecKHX CHCTeM) .
.[(mr
y.no6CTsa KOH-
Tpomr nonHMopcpY13Ma He TOJihKO caMHX CTPYKTYPHhiX reuos, uo 11 reHCTHY.JecKoro KOHTPOJTH KOHTpOJTMpyeMhiX HMH npO,LiyKTOB (6eJTKOB) KOHKpeTHhiX 6HOXHMllqecKHX peaKQI1H, f'CHeTHK0-6MOXHMMY.JeCKHe CMCTeMhi 6biJIH llOLI,paJLI,eJieHhi B CBH311 C l1X 6HOXHMHY.JeCKOH cpYHKQHeH. K HHM OTHOCI1JII1Ch TaKY!e: cpepMeHThi QY!KJia Kpe6ca -ManaTnerHJ(poreuma (MDH) (K.<l>. 1.1.1.37), MaJIMK-3H3MM (K.<l>.
L 1.1.40),
1.1.1.43),
rnyuTa
$epMeHT
(K.<l>.J.l.l.l ),
(ME)
HJOQHrpaTJ(er,L~poreHma (IDG) (K.<l>. 1.1.1.42); cpepMeHThi neu-
TOJocpoccpaTnoro
-
6-<Poc<PorniOKOHaTnerHJ(poreHaJa (6-PGD)
MeTa60JTI13Ma
neHQHHaMMHODCllTHJ(aJa
3K30feHHhiX (LAP)
cy6CTpaTOB
(K.<l>.3.4.l.l ),
(K.<l>.
3CTepaJa
rnyraMaTOK-
ca'lOaQerarrpancaMMHaJbi GOT (K.<l>.3.4.1.1 ).
B
OCHOBHOM HCriOJih30BaJII1 TPI1C-UHTpaTHhiH 6ycpep H cHc-
(6).
J1ccne,ll,OBaHHH npoBO,ll,H~1HCh cne.nyiOmHM o6pmoM. Ha nepBOM 3Tane 113Y'laJIH aKTHBHOCTh 11 riO)J.BI1)!(HOCTb pH,ll,a cpepMCHTOB B KOpiDiX, JIHCTbHX 11 nno,ll,ax ( ceMHHKax). OoToM cpaBHI1BaJII1 11x aKTI1BHOCTh 11 no.nBI1)!(HOCTh y pa>HhiX BH,UOB. CneKTp 3CTepm rrpeJ(cTaBJIHeT co6oH. rpynny cpepMeHTOB, npHHaJ(Jie)!(aUU1X K KJiaccy rHJ(pOJiaJ, KaTaJIH3HpyiOil.(HX rHJ(pOJIIB 3$HpHhiX CBH3ei1:. Y HCCJieJ(OBaHHhiX paCTCHHHX npaKTH'lCCKH Ka)!(J(all: Y.JaCTh paCTCHHJl: HMe-
HOCTYI 3CTCpa3HhiX 30H. 3cTepaJHh!H CDCKTp B KOpHHX HCCnCJ(OBaHHhiX o6pa3QOB pa3Hh!X Bll,ll,OB 3XHHaQeH npe,nCTaBneH .llBYMll: aHO,UHhiMH 30HaMH, KOTOph!e OTJIH'la;lHCb IIO 3JieKTpo$opeTI1'leCKOH nOJ(BH)!(HOCTH. bbiCTpilll 30Ha Bh!HBJieHa y yJKOJIHCTOH, MeJ(JieHHAA -
y 3XHHaQel1 rrypnypnoH:.
B
3XMHaQeH
KOpiDix pmHhiX BHJ(OB
3XHHaQeH o6Hapy)!(euo nannqrre KaTo.UHhiX Jon (y 3XHnaueH yJKOJIHCToii Ka-
angustifolia DC).
pacTeHHH, KOTOphre Bhrparu;usanH B ycnosHHx 10ra YKpaHHhi (XepcoHCKM o6n.).
B
B
JTa CBOH CllCUH$11'lCCKHe OC06CHlfOCTl1 llO KOnl14eCTBy H COOTHOUICHHIO aKTHB-
6eJIKOB C 113BCCTHOH 6HOXHMHY.JeCKOH cpyHKQHeH (2).
B
MHKpoKaMepax.
TeMy 6ycpepOB OopTepa
pa6oTe 6hiJia HCllOJlh30BaHa TeXHMKa rOp1130HTaJlhHOfO M11Kp03JieKTpO-
$ope3a B 13 % KpaXMaJlhHOM rene, C HCKOTOphiMH C06CTBeHHh!Mll MO,U11cpHKaQ11HMI1 ( 1 - 2, 6) C llOCJieJ(yiOIQHM rHCTOXHMH'leCKHM OKpaiiii1BaHHCM. 06pe3KH TKaHeH pa3HhiX OpraHOB paCTCHHH 6hiJIH OT06paHhl BO BpeMJl: 11X pOCTa. roMoreHaThl fOTOBI1JIH CJ1e,LIYIQIQI1M o6pmoM: KYCO'lKH MaCCOH no 0,5 r OTMhiBaJIH OXJia)!(L(eHHblM $H3110JIOfH'leCKHM paCTBOpOM, JaTeM fOMOfeHH3l1pOBaJIH c no6asneHHCM 2% nereprema Tpi1TOHa X-100. nepBH'lHb!H fOMoreHaT ueHTpn:cpympoBaJII1 npn: 1500 o6/MHH B TeY.JeHHe 3 MHH. Om6pauuhiH cynepuaTaHT XpaHI1JIH rrpH TeMrrepaType -20 °C B DJiaCTHKOBh!X npo6HpKax Tll!Ja Eppendorf. 3neKTpocpopeTHY.JecKoe pmneneHHe HJocpepMeHTOB BhlllOJIIDiJII1 B
To.nnaH Jona 11MeeT 6biCTpbiH H Me.nneHHhiH sapnam, a y rrypnypnoH. npncyTCTByeT TOJ!bKO 6hiCTph!H aJI,1eJibHhiH Bapnarn). 3CTepa3HbiH ClleKTp B JIHCThll:X 11CCJie,UOBaHHb!X BH,UOB 3XHHaQe11 npe,nCTaBJieH aHO,UHbMH ll KaTO,UHbMH 30HaMI1, HMeiOIU11MI1 pa3HYIQ 3JieKTpo$opeTH'leCK)'IO no,nBH)l(HOCTh. Ano.nnhre JOHhi y 3XHHar~eH yJKOJIHCToii noKaJaJIH 6onee 6.hiCTPYI0 flO,UBH)!(HOCTh, B OTJ111'1He OT o6pa3QOB 3XHHaQeH nypnypHOH, )' KOTOpOH npHCYTCTBYeT Me.UJieHHb!H BapHaHT. Oo KaTOJ(HbiM 30HaM y 3XI1HaUCH rryprrypuoH. BhiHBnen TOJihKO o.n11u 6biCTpi,JH sapnam, a y 3x11naue11 yJKoJII1CTOH J(Ba anneJI.hHhrx saprrama - 6b!CTphrH. 11 Me.nneHHhiH.
B
ceMHHKax 3XH-
Hauel1 $epMeHTHbiH cneKrp JCTepaJhi npeJ(cTasnen aHO.UHhiMI1 30HaMH, o.nnHaKOBbiMI1 no llO,UBI1)!(HOCTH y 060HX HCCJIC,UOBaHHhlX BHJ(OB.
B
o6ureM, no 3CTepmnoMy cneKTPY nonyqeun.hre .naHHhre noJBOJiliiOT
npe.nnonaraTh y o6onx BH.LIOB 3XHHaQHH TKaHecneurr$H'IHYIO 3KcnpeccHIO ue MeHee Tpex J10KYCOB 3CTepaJbi, O,UHH 113 KOTOpb!X 3KCIIpeccnpyeTCH TOJ!bKO B CeMJl:HKaX H ITO KOTOpOMY He 06Hapy)!(HBaCTCJl: MC)!(Bl1J(0Bb!X pa3JIHY.Jllll, BTOpOH - TOJibKO B JIHCTbHX, a TpeT11H - KaK B KOpHHX, TaK 11 B JIHCThll:X. 0pH'lCM Bl1J(OCIICU11cp11'1Hh!C OTJII1T.fHJl: na6JIIO,UaiOTCJl: ITO BTOpOMY H TpeTbeMy JIOKycy: riO BTOpOMY y 3XHHaQel1 nypnypHOH, B OTJIH'll1e OT 3XHHaiWH YJKOJIHCTHOH, npHcyTCTByeT He TOJ!bKO OhiCTpblll HO H MC.UJieHHbiH aJIJICJ!hHb!H BapHaHT, a riO TpeTheMy - H B KOpHHX, 11 B JIHCTbll:X y JXI1HaQCH nypnypHOH 3KCITpeCCHpyeTCH 6onee MeJ(JICHHhiH sapHaHT, qeM y 3XHnaueH yJKOJIHCTHOH. TaKI1M o6pa30M, MO)!(HO Q)!(H,UaTb, 'ITO BTOpOH H TpeTHH JIOK)'Chl 3CTepa3 HMeiOT Bb!pa)!(eHHbJe Me)!(B11.UOBbie OTJIH'lHH. J130UIITpaT,Uef'H,UporeHaJa IIpHHI1MaeT yqaCTI1C B MeTaOOJIHY.JeCK11X npoueccax QHKJia Kpe6ca. Y 3XI1Hauen nypnypnoH: 30Hbi aKTHBHOCTI1 cpepMenTa Bhl51BJ1CHhl BO BCCX HCCJIC]lOBaHHbiX opmHaX paCTCHHJl: (KopeHb, JIHCTM!, ITJI0-
105
104
Ex libris ТРАВЫ ЖИЗНИ: http://medherbarium.blogspot.com
-
I
.D,hi).
Y
Jx1maue11 Y3KOJ111CTOH rrp11cyrcTBI1e 30H lCD o6Hapy)l(I1Ba.rroch TOJlhKO
60JibllleH JJieKTpOQJOpen!'IeCKOH nO.[(BH)l(HOCTb!O y 3XI1Hauel1 nypnypHOH no
B KopHe. B Kopm!x JXI1Hauel1 BhrnBJieHa o.n,Ha 30Ha aKTI1BHOCTI1 <fJepMeHTa 1130-
cpaanemno c raKosoil JXHHau;eH Y3KOJJHCToi1. B
U11TpaT.nem.nporeHa3hi, 0.[(11HaKOBYIO 110 nO.[(BI1)l(HOCTI1 y 060HX BI1.[(0B. AKTHB-
JKcnpecci1JO aHO.[(HbiX 11 KaTO.[(HbiX 30H GOT y o6o11x BH.[(OB JXI1Hau;e11. TaKHM
HOCTb QJepMeHTa B KOpHl!X Bblllie y JXI1HaUel1 rryprrypHOH.
8
JII1CTMIX JXIiHaUeH
rmo.nax TaK)l(e Ha6JIJO.[(aJII1
o6pa3oM, no cHcTeMe GOT Ha6JJJO.[(aeTcH 3KcnpeccHl! .nsyx JJOKycos, npH'IeM a
rrypnypHoil o6Hapy)l(eHa o.nHa 3oHa <fJepMeHTa ICD. B nno.nax JXI1HaueH rryp-
KOpHHX 11 fiJJO.[(aX no 3JlCKTpOQJOpeTI11.feCKOM nO.[(BJ1)l(HOCTI1 aHO.[(Hb!X 30H 06Ha-
rrypHoil Ha6JJIO.[(aeTC}! JKCnpeCCHJI .[(BYX JJOKYCOB, I1p11'-!eM 0.[(11H 113 HI1X OKa3aJI-
pynn1Ba!OTCll Bb!pa)l(eHHbie Me)l(BH.[(OBbie OTJ1H'II1JI MeJK)J.y 3XHHaueei1: rrypnyp-
CH 110Jll1MOpQJHhiM 11 6b!J1 rrpe.[(CTaBJJeH .[(BYM}! aJIJ1eJJhHhiMI1 BapHaHTaMI1.
HOHH 3XHHau;eei1 Y3KOJJI1CTHoi1.
Ma.rrar.nem.nporeHa3a KaTa.rrl1311pyeT HAD-3aBI1CI1Moe OKHcJJeHI1e Ma.rrara. B Jli1CTbHX JKCrrpeCCHll JTOro QJepMeHTa Ha6J1IO.[(aJJaCb y 060HX BM.[(OB 3XI1HaUe11. f1p11 JTOM JKCnpeCCHH JTOro QJepMeHTa 06Hapy)l(HBaJiaCb B KOpHHX 11 CeMl!HKaX. B KOpHHX, JII1CThli:X 11 mw.nax pa3HhiX BI1.L(OB JX11aaue11 BhDIBJJeHa TOJJhKO o.naa <fJepMeHTHM 30Ha MDH. Tipw-IeM .[(JJH JII1CTbeB xapaKTepeH 6biCTpbitl a.rr-
Jib
HCCJJe.[(OBaHHbiX 'IaCTeH paCTeHHM 3XHHauen (KopeHh, JIHCTbll, nJJO)J.bl)
30Hhl aKT11BHOCTI1 QJepMeHTa 6-QJOCQJOrJJIOKOHaT.[(em.nporeHaJbi (6-PGD) o6Hapy)l(I1BaJII1Cb TOJlbKO B 11JlO)J.ax paCTeHHM. f1p11 3TOM Ha6JJIO.[(aJIH .[(Be 30Hbl y
JXI1Hai~e11 nypnypHOH. f1p11 aHaJIM3e 3JleKTpOQJOpeT111.feCKI1X Bap11aHTOB pa3Hb!X BI1.[(0B 3XI1HaQel1
neJJbHhiH aap11aHT y JX11aaue11 y3KOJJI1CTOH, Me.[(JieHHM 30Ha caoncTBeHHa .[(.!IH
Ha6JJIO.[(aJIH
y JXMHaue11 rrypnypHOH. B
(LAP). BbiHBJ.IeHbi 11 Me)l(BI1.L(OBbie OTJIM'II1H no no)J.BI1)l(HOCTH 30Hhi LAP y
KopHHX 11 11JlO.[(ax, Hao6opoT, 6biCTphiH aapHaHT
BhffiBJJeH y JXtumue11 nypnypHon, a Me.[(JieHHhiH y JXI1HaueH Y3KOJII1CTOH. Ta-
O.[(Hy
30HY
aKTI1BHOCTI1
no
JlOKycy
JJeHQMHaMHHOIIenTH.[(a3bi
JXImaueH nyprrypHoif 11 Y3KOJIHcro:H: B roMoreHaTax Kopaeif.
KI1M o6pa30M, 01.feBI1.[(HO, 'ITO B pa3HbiX TKaHl!X pacTeHHH JKCnpeCCHpyiOTCH
TaKHM
.[(Ba JIOKyca, no KOTOpblM o6Hapy)l(I1BaJOTCH Bb!pa)l(eHHbie Me)l(BI1.[(0Bbie pa3J111-
6110XI1MlflieCKHM
o6pa3oM,
a
o6meM,
'II1}! no JJieKTpOQJOpeT111.feCKOH nO)J.BJ1)l(HOCTI1 Me)l(.[(y 11CCJie.[(OBaHHbiMI1 BI1.[(aMH
3KCnpeCCI1JI Tpex JJOKYCOB B 3CTepa3HOM cneKTpe; .[(Byx -
3XHHOUe11.
MDH,
CHCTeMaM
no
Ha6JIIO.[(aJIJI
7-MH
HCCJJe.noaaHHhiM
TKaHecrreunQJH'IHb!e
ME, GOT; O.[(HOfO- 6PGD n LAP (cyMMapHO
13
reaerHKo-
OC06eHHOCTJI
B CIIeKTpax JCD,
JJOKYCOB). no 5-TH H3
Ma.rrHK-JH3HM CBll3hiBaeT rJJI1KOJJH3 c U11KJIOM Kpe6ca.
7-MH HCCJle)J.OBaHHbiX reHeTJIK0-6JIOXI1MJI1.feCKHX CJICTeM Ha6JJIO.[(aJlli Bbipa-
Y
)l(eJIHb!e Me)l(BJI.[(OBh!e OTJJJI'IMH no 3JieKTpOQJOpeTH'IeCKOH nO.[(BH)l(HOCTJI y
JXHHau;eu nyprrypHoil 30Hhi aKTI1BHOCTI1 ME BhrnBJJeHhi so acex 11CCJJe-
.noaaHHhiX o6pa3Qax (KopeHh, JII1CThH, 11J10.[(bl).
Y
JXHHaQeH yJKOJlHCTOM 30Hbl
.[(B)'X 6nn3Kopo.[(CTBeHHhlX BI1.L(OB Jxnuau;eH nypnypuoi1 H JXI1Houel1 Y3KOJII1-
aKTI1BHO(.,'TH ME o6Hapy)l(eHhi TOJJhKO B KOpH:Hx; B JIHCThHX pacTeHHH aKTHB-
cruoi1: ( CI1CTeMhl JCTepa3hi, MDH, ME, GOT, LAP). l13 3THX 5-TH CHCTeM
HOCTh He BbffiBJieHa.
MeJKBI1.[(0Bhie OTJJJI'IHH B IIJIO.[(ax 06Hapy)l(HBaJIHCb no 3JieKTpOQJOpeTn1.feCKOMY
Y
JXHHaUeH rrypnypHOM B J111CTbHX 06Hapy)l(HBaeTCH JKCIIpeCCI1}! aHO)J.HbiX
crreKTPY <fJepMeHTOB MDH, ME H GOT; a no cnCTeMe 3CTepa3 - TOJihKO B Kop-
11 KaTO.[(HhiX 30H ME. B KopHHX 11CCJJe.nyeMhiX BM.[(OB JXHHau;eu raK/Ke Ha6niO-
HHX H JJHCThHX, no cHcTeMe LAP Han6onee OT'Iermmo TaKne pa1JJH'InH Ha6JIIO-
.naeTcH JKCnpecCHH .[(B)'X JlOKyCOB JTOfO QJepMeHTa.
Y
JXHHaQeH nypnypHOM
.na.rrnch B fOMOfeHaTe KOpHeH.
JKcnpeccMl! "aHO.[(Horo" noKyca 6o11ee aKTHBHa, 'leM y JXMHau;eH Y3KOJIMcroi1.
OrpaHH'1eHHOCTh KOJII11.feCTBa HCcJie.nosaHHbiX paCTeHHH ne II03BOJJHeT yr-
06Hapy)l(MBaJOTCH Bb!pa)l(eHHb!e Me)l(BJ1.[(0Bbie OTJJM'IMH no JJieKTpOQJOpeT11'Ie-
aep)l(.[(aTh, '1TO BhDIBJieHHh!C 3JieKTpOQJOpeTH1.feCKHe BapHaHThl l!BAAIOTCH BI1.[(0-
CKOM no.[(BI1)l(HOCTM KaTOiJ.HOM 30Hhi (noKyc
ME2):
JXI1Hau;eH nypnypHall rrpe.n-
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2. fmnKo B. 11., Co:mHOB 11. A. feHeniKa H3ocpepMeHTOB )!(HBOTHbiX H pacreHHH.- K.: Ypo)!(aH, 1993.- 528 c. 3. Pe.n:bKHHa H. H., 51H6aes 10. A., CaMopo.n:os B. H. I1po6neMbi H.[(eHTH<l>HKaUIDI paCTHTeJihHOro MaTepHana pacreHHH po,na Echinaceae (L.) Moench II AKT)'aJihHbre BOI1p0Cbl pa3pa60TKH, I1pOH3BO.[(CTBa 11 llpHMeHemrn 11MMYH06HOJIOrl1'-leCKHX H cpapMau,eBTH~IecKHX rrperrapaTOB. qacTb 2.- Ycpa, 2000.- C. 66-67. 4. Baum B. R., Binns S. E., Amason J. T. The Echinacea germplasm project, DNA fingerprinting, and international symposium (June 3 - 5, 1999, RitzCarlton, USA). - 1999. - P. 187- 205. 5. Foster S. Echinacea: Nature's immune enhancer. - Rochester, Wermont, 1991.-150 p. 6. Harris H., Hopkinson D. Handbook of Enzyme Electrophoresis in Human Genetics. -Amsterdam: North-Holland Publ. Comp., 1976.- 680 p.
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Ex libris ТРАВЫ ЖИЗНИ: http://medherbarium.blogspot.com