งานวิจัยชิ้นใหม่ของโลก ตอน คืบกระดื๊บแล้วดูด by Sarakadee Magazine

runrn!:rrrrlrurhuoourDsrorss

' uunau[6,rn1 , <i'l ra'nrlnoarrruoorur,arld //tRii. r, nn'u Jnrn.,,.n..0,r,,u1 ",'n,",'o 3 / 1 4 ' h ' ; r = u c o u r r r . h n u r r a r J ! ru DrU. J r . ds.!ru:urirrarlounrnrnr,odranlr

Jii rirui rr]uut]ranlrnld

I a.||toirildorHrsJsurruurn nrn-rDldirra rlunrsoaau raa,nsDFio uir'rrurrl iouraoonnrluauiuiu!:rirldiJorir\) fl-urirrao r uoeor6aor?irldluns:rxr,IiO uuiinsoDLorour:arsorHrslS

ddooarsSLrauoonurrjus 3rudrn[0]a nul!l|naoorirDc-rHa\ijoorhrs nrnorrh.urar!rsnnr:uDouro! r uru b 16ouldiot]lrdoDauojsdn

nudouni uunari:rqiairdnuiourinurnrn ? rdcoucroonn )ru9rn,)runo)luufrsonduhrrr.rr. rnau ilnlsanbrniuo{iluc!rFu1 uiDdro

roaidilLiLiiudoUalu rdrlrnj orudrausj,n-uir nrnqulr_LluaoDsoolns uan!rnrnsoDriaciru'r Ia: rdruudoorodu.i4uriJrrnldd.uraLo rdu,r'jruu nL uua...urDns0nursrnuans dtrasaDiio:criia:ooruhnhilauniu

nrnnrsanisroD sDri'r nr!dsras!ro.i nrna,nislnurntu:-n!,nr.ros iun nrhiiirirursru.rnfarnharurtoruus farnh.rtrrri:inardi'rfjriirfi!r?jnntu lurtoaunsuasru rtu:i:irscnuunr.lurls:rnnjn! h anah6o{i .JaonuonourDuou b arsuusuoll Lau-anrnafa rtibtrahdetta ntilsr!1lrunisuulunru:unsun,q nionLturaiunsDrsnn.r;rnruulufiouuoDus:rnn|nu nrnanaHaelradlpsatia!ziroiitrrliharnharueo dJUs,aniiuu'iciJnnnuorrnunnooFoourr.irl]r Haenradipsa zeylanica a.u Haemadipsa svlvestis nuruurno'DrciuSrsrroounaroroDJsrnflaDld iounnaoa_r!rilunrnhlinhLjlusruDru urDarlJudaolhJ t oDIanIa,ofjs,Hjronrso:Diaan!rn]aad nrna uunrsns:nrurnur:nrDnr.l.i b urunrDnrASarura:a:iuoon. c ururnur-nrnrhar. d uunrDoaunarDroods: nfi

drnsslnsn'\)dtuoiunrdluio0din0ooniuooo_ruhu_l I jUUlUtaOgtan 1 0gclsoualgl!dthsut'11:6 u(,ltrhuorhdo !\)unrhuuronc|ann)nnoruusu mnnnhsoocY lunrnuroirio?:iurnsslnsri;D b aiu nlo[a]"criiu[0lagd A

rioD-lrnIiueodruhanCnlfi iJuhar-ntornrar-crhdo lrolhIcu8oorufiurnrDrunuo uu?ocoDasrD ruunaruruonnzroxsDurn rdqouooonuc,cHuD juqo&uhfi oicos:uuErurUrnrfl ra:idurolhqininr

n"lfi vs frnnn

-: l.r.i'trronlntdrrirnrn rLfi 1

rnuo';n:';u:rr.r rtri.rlriire r LihL{rrr I

A-

m.rfiillr a'lug1.! 11i0utull1 n1M1fN d0ldtuduotvndil

nrirrfiorrr{urio1]{-r.r a.run:rin rflu

utull Ltt

q i. -, x $ t l r"n u t l t l . r F l t 1 t L tuRl t 1 f il n I t i r f l

Lto:ouilrli nuFu-.r\riLo uL#l

u111

tLrirrrGLrrlnairr{trizurioonrrrra t n t t { n 1 ] | l l u l , \ L f l ! n L T ' t R o o t a 0 9 l :rrrhvrSorrilmrrnr6rrlr ra;orn-r lil0ttl.lt{nR!qnt[fl nrnauru tLrru-nil-untrr'hj rir.rriauaon"Lrl "r"rnrn" l.r"i'o -,. fir"hi'lfiriortllrufir,trio!n{urr0rn [[.i1.]-t11u ttlt{1.lt,u0.to t t0n1-l [o81aR"1 'huo-ooiu:" Fra.ifl l:aroal|al:Ja1ra na n-r1 ?a1n ro 1 nir frr:r-:rirLlr iradl,i on"s)o:'l: ? L{r:lrfr'l:u-nna fi41410j1ll91Rttt|litJn5:nta LtaYn1] t u ) t d 9 t -l R a i l t !

nirt

[l]d.t 11t01,!0lrlJ

tol{lilfitro!

n r d o ? l 9 r ' r l ) - : L ndfrlf J l l l r L : L Lu a n

{

.,"

! r . i ! o t n l f i t u o n o o l ] 1 1 5 t0L u t LLou n l

firrnrrrrrurirfr finrnorduoti" rrr aA,

lltntn(111'rurJ]frSL'r1 Llrtol]LL14lr.i11.l

n ! ) n u d 0 - t 1 j t o tl , l

{

" 110tl0lflul]0 lmln tl]l ifi Haemadrpsr

nYt]Y'lR lte'lRlll'lU.ilutnlI :illiltn0 1{U

drrdrnoraorhe nrru drduij:;v'ir r rJn,l''g v r.rnrI nrr LLa r ,i,.

nt5LUn81,](tU 0 tAl r t u L u S i l t n t i 0 l

dae fivrrlurir:Lna'lnotLa:nn.uo.r 1nflwtdutltLta:)-ouil ttd:tt.utn (tltc clc e) .]j0lntn lltntidRaln) dl

rJol"unr.rnrfl nrfl oia :strn:Trn

orrr:rao:Io:fl aooorr{n"i'ladro.r "aforihrinfinu...ftur]r,ur1nqj land leeches,lerrestrial leeches Ltei iorrrnrir Lrrfiortfrae'Lritfr ar:.rr:lr 'ldntrn]rloror:drirr';Golnrn" e tc, Lllrder rir.rnor6ooraro:or-nirnr rmlvqj1fi r(!:i < J ! 'l6ri-rsro! n { q : t l a . i 1 , { l t m n t { L t l na { d ! 1 1 1 L v t a ,Jra"o,uuun oor6ood'0,{riluorrr: rJSrgqr"bu'-n "ornl'-tiafiapirrir :r riruornu..tDrid Haemadjpsidae ,lrifrIn:- l | d 1 ] L n 0 :l n t t ! l D n t ot 1 , ] n t ! l n J- fn u a:riril"trn.trrv{oaar"r.urovrr ornrir I il! LTa{ lr fl a'1fi0 u Li s.r e . ! h, " q\Jqlnfr nooor.rd'r'lfiir finur lro6oroioGrxrlrln nt:riro16oo rxr6 tiLtl1]0:ar1tg "L00u nd!ul n)tl] L11! 11rnrtirnta l!,!qnM0dui!I

Rtul.lLtlotiaflu LuLlttutLlt)nI tu!I

LflA$O 1i?lrslrufina rlF.rurno 11 i . . - iq u d l 0 ] F I U o g t l nL u u t t t l u t ! 1 l ! n l 1 ] t n o

1rlrlo110iqql&!Rr-:nrfl oilio]l I L:01

r"nr'lri\firfi orr"u nroisrdri cr ntu:irotmaoi llnr fi aoril#nnnl rfil'ir nrnr,urlirdroionr:6nrrorii

rrrtfrooqjrrrn\driqifr,ir rilrr6o.r

"lri$irltn6rSoltnirudlorn "hri1nna:tua:"uhin"uron ":J:nE

'A A - - /A ot l:.ri1'!1rl11.r tRd1nllafll td0 arnd't11 nFrl6:r !u]d:Lllulln :lr]nnaa-r L r.l

, ! :"." 1r!1rntltulr alul L!Lo

t60nFtnttnfl 1141fl l\0fltllll0i0nnu

ar'loffnunzuuiLarurirnrn"

n:tond1ufia".r1]tnnii :]llttL lottn

! LFrl Fr:1'r11rrl .l,lAs |Ln:LL:Jn.r "LUL:JO

. a

". "

. ,s !-

v 5 0 ! n t u L t l t L d t l t t u i 1 1 d o :_ n d l ] !

D l . r:

1 1 t 1 , 1 d 0 1 r f ]n1ut lfnl ! 0 u [ 1 . rI1

t|q1l.1 tta!!Rs3fiflE

,,]a

tnaud9l1Yt:.1

tnr:o ullrio lrl nr,t3olr,r:r nr n.uo.r

iYc n.i Lu!qtu L5tffi1uln rr6ooaon; oirrrnmrnfrdr.'.riiooqi

ri': u,rr-r:o ulnddriro"

..' , t l l t l ] n 0 R L t ' U . ! vv:l l] ' t q n : i n ! f l I t n !

{

-q

t{ UO1lLtV.!

u';

LllUlL(116ltAltllRllULAU tllnUlrOnlt 1n.io1.lRl!i6lu tirsrou6-n{or:n rir"liil.r{odo.r nrn "rirfin" rLd:ao.rinnrr,rri*irfia or:lLrutfr rd'rotirrrrnorrnrnrir.r loouttlu0!nU niU:_1nU0l!t!Fltll[at 1 JI

tl,!b-tl:ol

flttln!1j:

aon [ai0Rfialat.i ^,",

d"r'

t1.r 0't!1n rntq ::!0n d !!:m1l a]a ')o111.1tfiu0 ]1L ll

rA,

1'64!nUL1llrflill1l1l1|ta "

.lJ L:D:UL)=]ll:

Fiou'ldtlniwia ,

R 0 t l 0 u 1)!n t t t 1 1l l l t F r l r o 0t d ) u

L1r LrILO rn lr,irnlYullrd:ordul r:J1 110o.t1,I1111 t1n1010{at

rdoooaluL,uioror:tidrrrfi adoua"

auo,f;u uurouu,flu"-'r''lun,5q1u uui nrr ni.iadir.rni!vranr iihrilr drd'! R01.11l1l1UltYn1!tO01llllt

, x !atl1tl!! nnnt:n:at].t0inn-]tur- a.t

ttt.lTtnL!toou lllu::LR11do)

t!0 llouq9tt: a{ulo lu}htu11l1.l}1}t1n q?:

11-Ll5:tfl ft--0!at'tt6

nullul

onil]111n04

!-rJ n1l flnHrL:ajl]Lnfl f,nf, rJ'r1 L149rLo

nx-rrdlii'!Vlt'r.! rfir!l6n{rainlrurrd

01$1tun1tr11t0goin0 0t. n1't

tin

mtn fl Lotta.l,!ti tltlitou Ll1tuu unll

rl tt -Lttdtn

i r l l l n L n l { l : l l r 1 l 1 0 l 1 r B 1 or 1 r L l r 1 ? t i*

i0ra6i'

arrnFj

aunrn rutritd nalt{rdrrrilLdr or6rsar :dtiuru.rar.^or-rr6oniirh-r... 'ki.utrur erlio!Tr rutnio r orruorRr: na?l]0nttnntn! L:.lt'tllLtu0tt{'tn8t0a

r l 5ARAKADE MEA c a z N E

\a)

io rar:-Lnfi o'rnrirr'lririra :rirnonr-:vl loso:.reiodrvrnvSo.ir r mrniils

olnlRfi 'lt|ntlltla01tan.l

6lnrrrrnrinuoil ra!nrnfiaR::111r L J A n 6 r ) - O n n l . r L n r L O l r : n fnl 1 : L n d l :

"tugnL(ln

L.UltLa'] eA4t

1,ll1R RO.i1.1U t l]111,16 U

t-o.r trirLtoli'-:tr#riu'Hrirvim"uiriull.r

orm::".r1fr sotLa:no"rnn Ln_unror'tioo 1 nir nnaan'l:l Lr';r:nr nl :'liLrar dou{rr!rtrnirn: rhra"lunrr_qo16oo ttn:tn]t1t:n -lltll.tll0.ittt LLn.llunol] t t n l : t 0 1 3 f i ) f i l ]l n R t ) d t 0 u l l l l d u o

a{11ln1!1.t111{n1fi :UllRln8RAU )

Lta't0a.idi1lo1.11l1 Llilt't.l0lllllllt

ulfr"

ullat

riuoonir LLa :r,:n".r tijlorrmra-n

| l l u 1 r u ] o 1 | n: : J $r : J o r r J n ! f r : 1 1 1 0 1 r ...olirn] nojdlr DEET olnnrr!on .un01nt4 t tlt11]fln1nt0ti.l oooLlJtutt1l

ortxdLviru lquadraj fil{dlri'rodr.r

t:rEnnrsLfl uLLrirr6noorrnfr r16oo

LLnuaL: flaaann4lu1l L1lr1a.r

rflull5:ar

a1]9lalr9jFlLaEoLoa

LL',: UOOO',t11l1d.l t11.t111UO tllt|1]! tna

A' (111U-tt1R|1ft141.t 1.11.11gfl n1j'ltA O1vl.l1

unnirfror: r:ri-r"liioranaroro6ou uiuqorfirdro'-:rr4qqrnrnndrodu .llan0nttn 16 L{run;rJ::arn-:rurirLEn Lilvdrv'lnd 1tr:14 tt 11t9to Y 111111.i 1u ''litRfi

.J 1nlln1 t1,{d1.1

ti:ln:lltd0

.id:

. u { n t l ] u L ' t aA1- } 1 n Mn 2 1!u- ,0 . i : u 0 0 u :i l l t 6

;u,

a.in'tnRt:tl.tu g1lg1LUoTul,{1.111 ufl1tu l,!1!0R n!fl!.i1lult'lRxqqm t! ntl:.r11110nou 0nnirno!! n-1r1.i t r t l 1 l t n n u o t t t n t o r L o : o r r i t o : r J

roidrro{Ioaordtrftr!fr Gsrod1J1]in n t 5 L n 5 f il n t 1 ] L i l l : s | u L d l ] L a o o i l 0 u l r l l

n5{R111fllt LOU',)- tfll0.! 111Ult Oi 9r0.i

v l o n S n t u o t L tnf il ] L o u -lll t n n t l ] t u l 0 I

firru"r ornL6ao'birirllnt:tnrs drrirqrriunrnnrLdlrJriir

frrrrotirrdu'hrrarrrnrrnr:6rir.r ornon:r.r{rd'rotjr{1tiJu6

ir:1ln

i-:f

1,I flaHrafi LIrLL-roaa:J1]ai1Jlflr I

l r l r J l l : n f l T M r f r : L : 1 1 1 f ln :

a ifl1l LoL1r0Iflut1rLtflInI

er-rriru10firLirlrLa:"[itt-raooon

tfl1:lti0un]l o-lutt?llooo11tlltaa1fl1 !4

ntd$rFrt0ufinr: 1n L LoLndiln

rr-rufr nurloir"pix6aou"r,!orl.rio llUlillllflnioOIa0O ruu riuwqnloo6odrun-r\rJ{r.rrir

:r-:rirla"lr]rulr"neu.rrrrno:.r{ru

Iooatd'ont:rn rotttto.r:.rtLtnr:ar

LLfi.r 3!riruriro;r'o.irirroon'll:rr t" -.

-i

ft1.lul]:Rt:ltRm0ntmt'ttlt]toun tnan t R ac l n : L o t u ! r d l o t n

iu,

n1911._.uuLli1lltt11'ttvuo tLtuo9rfl1.t1

rlqwinr:rinriorLfitrr LLrii6'hirrirnn lvninirorrdrll

riorirlnXuornooil rsur,liRrfij nx'r.! n::o.rirrrir 6r.l:i nurll trirr f l : : r : o n L R ! t L R 1 ' !l1l R 0 t n l d

, v4

tlru .i tolllmo

l'11{Lis.rtL'iuo o

6"Lrirr onnL:oritorrir6o nLihotir.r

t L ) t n t R n S L 1 l t n L n u L t l u 0 d g trrird-r 1

m l 0 l - l u o l : . 1t l l - l t n I f t t l l u t 1 t DI l Y u D

(|na;t9r01llDitn!10:ln tl1fr0.t

fi jtdoarnarr-r:]u1r11rL Loa:an -

n )-0lntdll.l ltnntnnl5FInEtR0ltvut

i nrno:oouauor otirrrrnsionrrr:r L1

l:dnriri'noo16oo loro:1{Lrnn"nlnr! oonfrrruilaviirrrar'lrir1lrnarrdr "lunr:dr:rlrrrirLa li.roti...orn

aud:L

natinut|a:LttuoFt0!9]nt

1.r11,!'lFrfl't! u0nn1R!

nifl 0.t

taonL:Lltrrt'll]a.i L9r D.lgln fla]q a.l

tRliou[nl

f,{ ,

00ll

!ltt1t.t1.1t0uo.i$fl1illno

f l [ n 1 r t q r 1 F ] 1 4 0 0 1 1 . { 111unur L : J a n :

{ au' t ttag1,ln1an5.i:]u0 rn1 L! 'tt1.xR !<!

,J^

, -,

n': e'l'ttryonvn!fl ut\1tlt tilluR1lo11n

n.ln vs III'I

arLla:LL6.i Loa !n::l|5fi.r

nr:aoaifla.ia11:.r11 rJLLa.rvianJ

m:.r

!!u.il!tn.t

a t i l ! t n t 1 l t : u : t n d1 ' b r r n r l l D o l ] t r ] l ! ",

9DntSdtfttll0no)0 lltR tt.lto00n tnu

lrFioo!Rarii!narrdudlfirrarioofi o0ll1l1fl1l11!t]:n: uu!niu0ilim1]t

t 1 ! l u . l l . )J 1 t 0 1 t 1 1 0il 1 tt t t n ! ( 1 i l

rtarorol{rarsi6r!a!d'u

t\qflfl::ut4llutttnmt1]o

'1rorv'Iornrnrnr:odrrfi rqrn"h"

rhrrrhn!rrrrai]]drugr urnr:l R 0 u u l . t l ] l 1 ! 1. _ r x l f i l u a u 1 ,l]0 u t o u l

rx:Jl:rl qqrnoo$voL1ru0uJ 1];or

i l ! n r 0 ! n t : o 1 1 1 1 1 t o 1 1 0 a r| ltnt t H 1 1 , 1

1 ] n a l ] n ' l \ o 1ttu n l : t a 0 R o 1 t L1f i] n T ca

0 o R e R a t ' t a 3 n 9 n L t o fal n n ! R l l ! 1 n L ? U n i L l ] U t n 5 t O0 n 1 t "L t t t 1 r ( t 1 d i l ,xa-el 1llllT llllnr-itlluoLout:-tttl1lln:-ll

vrtfrororaoor"lil ,

@o ]rl.Jl1l

LLrnft arn'irL6rLds6n ,," ,r.,,_,Li.lu1tn ":otuudu n1].t]1filufl

g [qidolitlartllto :ro15rua:fi rnr^:rir

1flan$:JLLn:

nl0l!

lririooariuir nrn4orr6oon"ro:"L-r1.r

, , ! i" . { ! fl 0 lotuLln1l1t11duFldl,,{ufi : ntnu!

nlnt

t|aiaolnl1ltnn3ltlluo

ltsrlL11aluLLTn:11 Lri!

'hrir

L B r 0 g t R d R l J lJ t11LR1:LOfl5r:Ol ltl!OO1A.i{!.ilrtU]0

:1]n't!lt'tnuR Io']1l!un$t!do'l1i1

-Ad

nila!\loLuoLli1ra11ltlln!

n o rr s r r

arrnd

11V

110tu:LL60.i

o3u! L rln:6sr'rrns.ran0lirrlr

': .i t3on! 6ornn!vfi:oonrRRr:ornrilu nrl nir.rroat|iroo1r6-.i]i5s:JTfu rlt fl rJrr11t f] Lult 11.1t?r ilrit o L(1j11111{ann 6ar r:nrir'161flu1 flru *eifirtihfirn r11ir'orrn r{riil(rdolroro{fu rn, b-$ fraAurn:--rrnRnr.r c,

-.1

"t1lqn9tI.l t1]1]aattounqaolulllJnmro0

tflnl11,Jlln0{!u11]n1ulq tlifll

"fro'irnrnl:Lnr:lotr'hir6onir q

ir gntl1]lrr.r,lruurn.l Iu1.1d9l )111o t1,1u

x ) e rA 6l: r1,1U:Ln1VU11l

ddru rnrih.uurn

nmfilrflrli

fioor6on6!LL6': lotlLn6srdrl:"lrao a:rl'irrrn Lihaorunr:nirir'l:l " s l L n r a r u. l o n l d s l l f l n t q l l L a 0 l n I d o

naliniMo tLi! 14lJ111.11[ul]rqo[o1tuloiLduoi IqiLioll{trLr]:uirn:tfi rl q

! !. 0|1u1u't 10vI01!aa.u{a Lot:0.1}11n

fltDLtot9tl4 6r tt6: Oo ryloqlflddtYlu

rfll.t0ltft1.J to"!.1n6{n1utt1! el ngl!!11

i n:ru{ld.rtrr rr nirn::rrr 6ro.rnil

)

,q v.

u J ,i L l ] L A n . i O A ' 1 0 f i 1lrJ i n ' t 1 1 9 1 " U O .!rnt

rnrnrs -irdarinqLr'oirrri!3i1 ua:rr rarrrurndl6rrrirrfu !?

rtuvrrnLr6unirontro:d'.rtno

qnrfirnir rrnruun1tlqjJtnd

16oo'[ir!16s!6i.ior]o"rlrnsrn'bi

0'1ffg0 g In$ll3llll1,l1loll

fl10tLotnYlq0.iLuott9ln:00s:

L li'ilfl {a1u "L Ut : . 1 1 1 t 0 If lt : l r x 1 t l l 1 . t d 1 f l f i t ! t l ] t l n ^e

:,e.l

$11.tt!0ntdvllll!40u00n

l)-d19tl

o !!fro rlrF!n nt t [1,,1t] fr-:r l! t"ui.r 10.1L:.r L6aqrSoar:arfl r:hn::rr,!-r: ttfil

'kinnernoorrLio.rlrR rirliiL6oo n R r t L f l l ol l . t ut a L l R l t l l t l ! t n 0 f l d l l i rnon*r4rar.r ari'oLorriroanornn"rriuLor (01l.i11) LtdrLoonrnru"h\ri {o !li!!{1Rr:rvrl:"16:11n Iofl i.r r.rf,si::riror:rdrr5n:rhuro a-rr:"otoo

"*t

tllo5 Itiaiti1]nofltl,Jl!ott3lni

0llu

l|a a

|l|ln:

ntl

11.ldu1l116

lflrJlnlnliunt

mo nO

tu,u", du ttoin;torirrnilrl:grrflr {uii11rl1l[a:oi1.l1o,uonrrn rrnn"r"LrurauloLiltrirqr ttr 6n:1fr roirotir.rnrnldn.:fi 116ontfi o q "

he"

f l a u n l l , { Rn i L n t o n t S t t i l l t 1 tt t d d do L dStBUfl LtU! 2--,....! -OOn t1L:',l18nn1|]All tAAX!-)t tflO

finaroiotl alrq:.r9r1o0'ln-ln1lltunll systematrc sampxng

llllilllr\ llll'u ul9l

'i11r'11.r oo no 0rr.lrarr o 0111.i!1.lfi: 1 l r . r L o u 1 r a . r 6 a 1 L 1 t u t : i t : t 4 15l J : l t t l l -" ;41 r' - - , Y 6 nlaLu9l: .i 1u111tdlt11tlL9llJ Jd-l1.Jl

ii'.iRar'l

N']fl VS UAD

or"rL6on6.r"lvaoonornritriu rnttra fi LtorLnr:ldrioluwru d'rdra'r6ru tilu::svLrarrnurn"rornnrnooL6oo rirnra rirS'otrlotirr rd?nuHr u]o[,!an:s'{fi ldoolva'l6n tr-kouartrvtinf A--.1 rl::rl !1tflauom114q14!1n!u1!qorlarttr1]i oo tr]r4n.iq0 rllnl]o-llllu LLflnnlrL Anfl LlrlUJrO)llfinLnl:slllLo !u ul0 L5 }]1:0nr.._ni]jaautnuFtl tnll

R011tvln1l1141',t14111fl tiAOflUflAflufl i

fid'u,uo.i 1nllararfioqo(frr) Ltdrl:\rialrrir6uniriirlr:nu ir.r v ?.!tr u,4i n : l L O U t O d I t l . t 0 Ut r ] f t t l l ] n d t t l

riotroo vtfr tLqidr ntutilunu rrir t6oo'lri'l6i a o n r l 0 1 ! ' t t r l r 0 R 9I r no r n o

d11.i1!qoRll!n1:n!o'rrr56nsnrvu

1]rn::o11r14i$enl]RoutnLLha L't Lg0

n15|LU.lO11o{ta0000tu Lo r 0. ln a t t] i

Ln,) 11fl:n1:nu4na.r Lllnaln.i oo-5o 1 1 1 1t1v a a 0 l n t o ! L U n n R u o l : v t o

n u f l r n n : n : 1 | 1 t o L L l 1 If l tu n l s g n I L s

aar.iYllnof! ar]LLarrni:tlaou

d3'rinli!rirlrnrliiriu'lfi ,

-t^

, x

" - --:

n-t'tun-t'tw0.iIt1]oovar!o3n'r1 R-nurr ni-:11!i9txd.in1.tolt[a:tiallai!l nt]t]nSlwo{|t'iuonfin"ni5: lflr o b rldd{l.lgll--1]-lntan

. )+, v .

Liaorirlii ltlriroornr:Ltiln'r.lar 16oorqo\riaoorltro rE'r nrir-:'ldirnrn\ddr16ooqnruo tu[tJ0urrf]r]nrnrr I r Dn!d n:

.Un}\t1lR:',tO11:tUtljydu0q tn [1rnr14l0nrL]RnqirYlro01.l n114AUa{(|n?!1UllR I ,," , { f,. 1 tllln u" ului1.llll.l1t Llilot!:t1l ut

t1ll1'1lliIn ltu

,irutJ1n.i!11.11!n 5{uusLvol]lR

ttfldnoldo n ]tfl i_:o1u1to.i111R -A LL'l-rr14rnr1fl 1 loL :Jl:n1r'llfln { l g R 1 1 l l ' , l 1 C t ! g n !f 0r 0 3 o1 r i o 1 o 1

6 l]aat1l0i nd[11]1lln

Frr|La:flr:s]1rL1,1Lnutnao1]0t $tu01atg

firirn:air.mrrultoirs"lii:iuroqour.r

rii!rntu,r4nr:a'o

iru"uooL6ooL:rtlni{ostr'l:I 1tur1l',tn4ot6oonr aogdlsrni 6 r6lr (haemadin) oonurlirttu r rrnuna firlorarLridirl{L6oo'lriri

ur m:#rurlrdmrulrrh LfioLGtrrtfr

rrotLironra"r (Iourii::J-rfulilrn"!)

,qo1]ln:0qI1] fl r I "tllrlrlf rurddofl111fl .u!u ttflnl5llulll1tt1vilnnalfl n:tion rLa:rmoo6oo"lu$:rrgd

"nnril6rr" LLrJ rLillaornriaoirr lrrornrnloofilrrrinrniiou',riroutur-o "Lnoarurr-n 1 ntn l)!l1 l)o0olllRl!

L $ntma _fltntll Lt! LUlnuailuonlj

tLogtnr:i.i'hiLn6o!'l rrilutrarrrr! ;, .u.rnlno1tl.rrll I n:dr 1:-nlnT:!.i0q

_u1.lLll! tn! tufi n1 L0n d1]olllt)-D tu lll ind "l {tdttnll!:

[!tnn01,]111-)Li]1lL)ar

rru (Lwr-r:tndoliiturn) or:.ttoeao.r

rnu I rihr:ur{!n1u1l!d.i b 1601r'16

'llrirur:rfi trRr'r''lnr o oiurir1fiuna i

nr:dnrr di qrq'rr rlLiuod oaooou

mnRLnt:tllultifllonllt

-.A"-","qqntdl0ldol1ltttflld l 10u $t r!tr

ni",uuun,n#i.rlorr'ln, rr uiu

" tfl S]]nn]]nt:6n!19ll1.1 0ut.tui11n:qei

oun5 ifi1r rSodn!],nfio Gpecies) 10.iilo101u Lofl1:ou nlngD1rt1fl.in1l

nt v'lI uI u"ld91"'? FrFl

|l.]lhFD v !

i.,

Lr.r1flun.u0.!0 0l.l0 9tlll'l lR I i n t g

nirnrro 6rLfirrl6rtnr:rdr rar usi

nt't.!il-t'tn o.! [sr tl srt0 rl, u fl:-Ju 0 0 R

1.""r

I't1t!11.1$\8nl,ta L!0nttnnDLoulll L1J

" r.rirlj:uu::l|u ryr! rl0ill!Y0tlrudun! "ro Lirrlfi ritrldooarnr rc'u ua rrrnroroL{r1ofi o6nirrilldo'i6ryrn "r{ rnr:'lRfi rdurrnr:rrorn"anritLn:nrit 9 0 ' l n ! 0 u n ] n f i a 0 n l n ! 0 d ! t n t t l

dnrr6uiiotiords riir6ifl lou\ i

tirl:rgridri flo

M l u i L ut n u Y nn t i F I n ! t / L u l l l a 4 a

firio.r i

LroYGnr r a ! t t g n l l _ l r l n r l j d D u r bl Er r { i rI D r n

!f l d l l t 9 l l t d l u t t u u 0 R n t n t ] ! g i l t u t"u

tuitirrl::luvi 6rsrdnaS::rrrnrcrRrw riR

ooGilrnarit|6:or;n11d6ndrfl n15:tu:tulto

ntnutRl]0!ntl! , i" < n . F I .6 c l d b ' v l u l t m ) L a n l . ]

LLsr n rir.:riuadt Lrillfrdo r-:lir.: I rno o .1

i urfi afli iruun rnofl16ofl Lih6'n-i t n t n l "lrifin::qna'ura"roioa{luh,,lair en". nrnoli 6)stnqa dd rfiq fidrddr

ourfifl 1 loulY t111]dnsfu:tu!||r.t!ul1

-;, t r d f -, - R t ! t d r o u l 4 o n u t"ue L- , t" l" L l l u t t a l s r ) r l M J

ogffiu; Nrtordrn"r ua:reJ:rnn:arfi on

ioi:.rit{ihrlfioi Iouflrnf ilurvrldoi ,

vlltltLlldLfi

ttul.llitno

Ltd:l.ld01Lt!ff

1 n d O"L 1 r l 1 r r l A a 6 r 9 r 5 L Al ifdJ

lio (d dq;) rorLot4so;iuoon Rt:o! ril,116:or1 rirr.rr:rlir a nio (a rAa

dla) ollJdl.lllrr.rlrnu$nlrn]r 6ra

"ltrri'rr6o:niurLoirarlliuiilud'rto.r'lu"ld ata110lLtfi nn110u0 tFIa!aan'11 Lmn lcifrns.irllRri !!r]R6i d rio

ririr fior driou{rrdudor fir rrrqdirronLrJrir r:rfr lirdrn-rrrn

i" t 1 1 l l r ( 1 4 f l l r J l { 1 r 1 1 ! f llrq n u o F l a r

nnt0-l.tLtlt-llttdn 6i 1l|o

HaenadtPsr sY/vesais LIUUL ilr

oiofirLIosfirllo{nrnriiro-oo;rfr !il f .i{rrru'lidl!frn:!'l1JrorFnId: !:lllE:Jtn.u1ltxudtoll

611tfl lllt!q9

r: otirior'l:lr'rr.idrr,lrh !rl o{ iir !'t''r 1.!lllilU!!1i-lin0[

1 . ]6

L L14Ml LLA1ntJl

zeylanica

lltu:

Family Haemadipsidae

DtntFI.Udl1i:3t1lFIn1Ut00At!:n Ll1LnO

-<-

n l n n { t l l l l O l O l . r l r iT l nan6:i

Ltnlnl

tllnflItula:L:ll]outotrl

iiru,uorriro:riluLriurnnLiluorrrrirniootir.r'l:66 nr.'rdulooldn nfiol rl::'iro"':LLiuooyrirfi rhn filrnm"ln: â&#x201A;¬rud11,l lLlJhl]nnnoovl: llr Lo rrnr::l|5 ,

lJi:RolJfr':rtr{lt!ldn 1 rtfl.in! o tt6]-l lnirllfiro

l|rn::Lfr: lo au rL9llrvrll

1160.!(segment)t]n]

1ri6or1r.rfirlood

a'ngnr:lol6!ot{0fr fiorrrr.rornnata rldor Inorli'ouirrunarr&o"rrl::no:r riol{rrlnn LLo;drzua"l#inrrlnorn fi:o:ldordotr(annulus) r1'rrfjm ob

n01litalnfllt uonn1nlj0 at1nM11ut

llou[1.] "l !41!ato-tdr!14rrrd:

SubordcrHirudiDifomes

"n-ns g t.rai :tt dttn:Rfi

t : u u . u l j f | u 1 d l L d u . xl R t f l l l l l i d d l n i

Order Arhynchobdellida

n:;o'rooq'lurirrrulori-r.:

!fi'o.r.irer 1 diroor"or:rrrotir.rrrirr

Sub.lassEuhirudinea

4 i.t , 1] tli:gtllfl 11t111l lt:tu-lttxl ,-l u tuL:Dlltln0 gRYuft ttnatfiton:ut{t

SubphylumClllellata

otifitisruiurr'6n rilr i 6oinor.iiirer-') 1Jatt]14tdfl 1)fil.ltR tLonnl]1]ttu u .u0.r11fl os rsno!]6'18:.rdou{i'.irroi n1rfi1l1,!!1rlRrfiui b rio hnqnr6flr (t11111r L1r19r:lrAdrLnA n oro1r dni'rilrnira'ntn r6(il

i::urir riruLih,uo.roir roI

4.-A

rLaz

nrtfin!rfi'td1ir:.rliiourrir'l \dn:o!

a0.r tltr

ur.iriffrihrfin.!r.ido0.uo.i ;ioisir!"1r LtqiorrJf, e.r1uoa rl6or6rfi.irurr 1.!tt11:UtlnRll!tn

Haemadipsa

nr:f,nsrrn.roun:li u -.-'

aon dllRll11tflo1011

lfl ntnuiL'ifl i

1 0dt.!t19rLt':Lglutsrr11r9l99r!r.M

Litrrnir "mn" loinulllt ur

-t

6g!!l al

LX.,.s.rrrol110!n!!4.!1,]'tno i!16dtrI

r]fra.riLLil4o (sucker)oi]rrjdruxil Ia:n]u1r1u10.rn1tt 1]1l.tul1tuoatttnfl nr16!nol!riIi!'lridr!rrnnr:1oo'ld lnoJ:n6nrnr:Lnr:fi rodnfu id:uuirt qoollll1l!41fl1

t 0!rl,IuoHl11r.r1 n:

Ln6oufr IosE,ndro-rorraon'LlouaoLL6'r larL11.r99l9lr1r.1 rooirnflon!

1ln ) l't nln

llur'sldorLLiuoofr ru{raaonun:6.:ritn-r llrftlrdrloafirou'iuoqfi u drmLnr:!'o rldooLLiuqofi runbrLa:ioriro-rlioroon ul.!l .tut {0tn00!11 L!11 11lril0Dtl

riorriqrroiLmdil!$iu6rrad riorrilnlo.rrtoddu{u{rrario 6r dor

rlf,a.rr

or'or:irrtri

or'u-::ilulilrf

111111111R

,t1 (€ye)olur:d$iriurLdr Liilne{ir $mlr|ler l]6nurutrL'luq'rx'rrlurr nrnr .ituru c o{Lniouilqodrllrir Liui6' 1.'l rinurd'{L'idru#rildrd1! { rrillgndrllrnr (anreriorsucker)i nror o.rtiri'r Ina1ttLLdn:ndarn:n{ril ffi drrnrir llll1'lr6nn'i1rL'i n'd nir n1d1llLL'ixnnnrt:llintrl:l u { .qjdr!! .d Lrlr drlrirr d " riur:liuilnndrr#rnn6nn-rft rlri.:lo.r nrrYrrar)uganrulint('uic|er an!fl3 LUn{rl|nlu a.n'r-ro { Lnoi:oq!Lrlrl r{u oodl!xft alin:ilttros1:tunlnLnlnu tiulnro{nrallrio.:rJrnn-nr:qdruft rlri.r nrU 1rn''tn'Um|*,.y*:.,:"".1"1'' uiv4ofirwa'r {p6t€rio. sucke.rr;a 01lt11.l 1.rfi?dr$rnlrnnrll o alr aqfluuno.$nn caldalsucker)!n!Lui!na!1 lltnolnqj (segmeno,illrllr rh:nordrorldo.r 6t rtnluull:tln8nln0n6nrorrnuru! tunnm'iiiiliLriuida.'a''r'fi fir Jar! nl!!!n3 |1rfi'' Ln'L|n3:alri:'r lr'dLan 'raN na rlJau l'$rlfr 1 olxr: i6o.!1rfl1!6'rldlrd! (hermaen6r1Y3rilor ,fil:llrn6o.:don-lLirtLnrLdtr nr1fi l n n f i ! 9 n n n ! f l l f i r . r r n a I L x 0 0 f i) 0 r : ! ! ptuodrtetttnr{n! u{lufllloJl!!n flo{ vn::1n:in"ft fl :nfi un"rdutdosrllnrlli qqq1n1f, riaon'li i"o"ooO nrulu nonlin:i lda{ e-oo r,{o.rrn-rnlRefl16oqnrR tlrtlotLilutYtlltu lr-d tlllolll tllRfl0q l1r10lnrq !q1| Lorfll rd!r!qrrlIh]

n:r1r'li6Mu .-d lonu'll rririu rl::lrt! t"" iu ri'rriolrc{n Lihn"ri.:i v

" 1llru 0rl nulnunrd tllllrfltln!nu

n:lnlRn?ttu

nll:nfn ,,llYll.ltl'llttRnll'ln[fl61 0en|606 ltlt]t

(anu)

ninrirrr:rorraadrrnn-orilugrJth!ron {pl nl Y) flr (teerh)ii6r0rior{iruxn! rfllriJ io{o : 6.r.t'o$lirlrdilrfi d'r!(xdollnlr]idn tTinln : Keegs, H. L., Toshioka, S. (secnen, rjdo.r i6u1!nid od Ji6r md Suzuki. H-'1968. Blood suckine Ioulilln'irLddltrd1$!6iLrl11inA'r rdr U Asitu LMhes of Fnily Eimdidae atd udnv:lfio.r:h:nor:#rurldo.rtiou o-a rr EaeDadipsidae Bio-medicll Reports of rif i:rilrrriAnJrrolJfia{ (sgm€ntd the 406th Medical Laboratory No.l6

oifil]urllit1$d1lrrirl

.!o.rdrn"1 a6ni!M!enft l]mij

n'n6''

Frrra

Jro'l

lozor-rLLrin:aYro,,iorndrail "o"oo. fflu11ll]0n01]!antt1]mSnnR!lirromo uirrl,lll6n 1 drdrd:vlrrnr .irrrlrrn

1]orr1ll

rnnrt tn It rit0m:tlrdln!rd9l)

d-oo

110 lod t)lllltllllm

b,,

, o

d'ornludx:larr qorduoir i

[ntoolln1ru-]tfl n.!$tqfinR]g1.lfl ml]vnJ d-llltvqln: .uu-lqtnn |t9]l'\ilnrt jvln

l,t0l

ann-111.t:0ttntl t1,!:1:-tns-lqnuui r-orlonl:LLqnoonLri6otL9i "Inol"

'lnf ionoaolflorufi ariir,uo.r:J6r m:{ihuidc!nfi rsrfro:rfrriou{r.iufi r

rrn"hifi dola.r(aififl!nr!roRnrn 11ln ua:rlro"hnir-rn,u'r.r rrdo.iirrnit,r 'i')u0u t1.1tl,l91fl t:trduo6a0l1,la tfl n)-n o"uuouu-o'iriluuuru"uono',ouuuf; a "lrri1 frLfrnil"lurir.rnar.rq4nhinr: v a t g u 1 1 1 n ( T 1Lru ! f l l 0 ! 4 0 L a S n ,tt t000 1tL-)nl]nd!u nll! Ulul LliruLfirllorfiu ir"ld rirnrrritrutfrarrirrornrr xtrl1lo!nrrllt19rir0nqo n160!n: dxqq r fniroraran56onir : -u,.1 , , Y LL6v('r.i f]]l FJll1rfiilr1l "L1lL]!U

dug tllt.llll t1.tllult fl tn 80]l to 0!n nt 8

:lnnqrudroloar3oLau1rl"Lri nrn

rih::u:

Llt1ll1liR ltfU: AlnnO.iltAiUtnn!L!1.1

-4 9 l 1 1 . t U . l 6 1 1 1 1 l n ALA1nL O n l l l a t

v'tn

dririr{noorrr:-rfotjr.lrirornlo nn

Ioqu'hdrrnarq4 r nrc'h

--g-,

" unntJflu 1.lRa0{tl!tl.t1fto51ul0

lrqj Lril {rr rdon3onfi\ri\ff rrn

o-d

6uoir"tactivephase)1]or . !

dxlrltj0f[]1Fr0rlql[n{n.i[nto0u durrnr.rirnlnrviliidodrltornrnfi

bo 11! n"-:tiaul:i[n oon

ntnL! tulltol,\0.l RULdllot! dtn-)

1'!tiLlidullLUdR

tnl]1]to Ll.tt!udod t1t

or r_:mru c-rt fraano: aiun:oonm iooarurndru"biqjrfl rwrorEnua: llronnr.r rn:firr nfrrfirilRnrn\ri 'ir!r riau rn ririfirisnir:ss: uronnrr nr._niqtdfoto$rrnlrii phase)nrnnrrr' nrtuJa'olpJfr 6onir metamoryhosis'?qoi.r" (dormant -, ig (rofror'lrhr 9t-l00u Lno:Jl nnu t1lll0uFt-lt9tll anrfl r:a siorrnorlfr 0t ttfldDnYt tqo u n1lt9ltu111ltfl

n']n vs uju

tultnuS:fl tl

zualfinrndru1rlqjLir{nrr:'irda air

Ltarll,lqotLnmln0iLlllfl

n i 0 0 1 l . lc l l l l l 1 n l ! . i ( | 4 3 $ ! n r : n o !

ir'hJLrfi r rrarqnqrnioriorrn nrr:nr:,irdn uqqdrroriri'or'l tlr6r nrnLrriruirlo:aanrr rarn'riuf ra:rrr'hi -.f

t o u n d l d 0 t a g t t gt v 5 0 T t n 0 ) f iu l , l ,*

-A!

" "14n]600flu(aoodflnntJ 1lnn-r6rrmn:

(Lriulrlirqn L:rdrnruLdoriol; u,

011r nt!1n0flf.iraonto nto8't.lt: e

o'.ri'u iiirirln4urownnnrqlrl'16' r'urylo 6io;cr!r:ooti:oq'liludl qqil!!r"otild irfl n uirflr:rodur!6iLfrrsl"ro thi'lii o ''ornnr:6nur1orqir.r nr,trw-dr.iLrnrn:.rnru ::trg'l inonluL6oufr e

ri:?qroutrlrrnriu

'irdal gdoiinr:agalorfi r:'iiour.i

Lltltl0nRtoaul

ih,4

I a:l nn nn1 fl:.]|Y0r11a.i4ora09](axl!n: dorrrr,lrjlv'hi{r nr:oort6oorirr:6o

LFr)Itil notunu!fitudtnt!1o tYl9l

6 1 1 0 r 9 l 0 a l r r : 1IF f rr b r l n r . u l ( n l r :

qq(la{ 1R,!u1o(an(|ailut9lRal.im

Lo:l n t 5filr! D.!I n 01,]Llto t l

'li\d4o16oo o-.ro.r fir qFnr'-r1tlnr: !1

o0n1l]01t5 an600na0lfljl ant5 <

r.ii'rr" ra'inii'!nrRllr"baWoi ri'r

Y {J.

fi ngtFrl l!11 [!tlqLta rfl !mrn(ll1,]rqldR

rLn:rutonatrltta'odrlfi arnit,uuro oaonn3lflLr]rrirlritn:rgliadri r rdcrr!ir t,rg.i IoirVnoonlrn\dlun'i{,1''l )Y ^ L9rlrlU tnn:nudll.nSlttdt dnll

, v ,q duln )t lul t 1o!qo itun lIo to011

arur:nrnorm:'16orlo:o'rtr'l hd4o"

ra"lornnrnqt-r t6r-:-on_uornt:ni.ri d fiurn:r6uqumnrrrnfirxdirr 1n rl::u'roru-a d'rlorririrfrn:oan'hi ! !rn,u0!a LLf o:qnuon\i\1116'drunr:rauril{in'f|t1J0 ulr'ln $.ru.t01n'1lo1tn , tr i: niror"r{tarfr orruS.rduiugrlir'lu1u LLfla:Lo0ht!1ri1nrL5:J1'11rfltn1]111tltn fiowolnrnri':Lis'tdirnr:zuarotiriou olR tljt!t9t0!1,!q!nla I Y ,a rufir uoilr6oofr$dn6rnrruar.rliai 6',ltM'!.iRal.rqqr{!o.rt|nloou ri.rr:.r r,rYo:r r; w6nri::tr n:vr n fiqrrouo'riwrru o:rrnrnfrrfirdn siol lrn'ldLto:rrnruroL6nLillrirlrvun i<' h i r o r n r n l : : ^ r u o t ia" h o n t ! 0 t Ygt ! n ! ! ! t 9 t n n I t t L n : ! u t o t 1 . t E

.! 10un1t:tllroLLdoItl.rL'i:

fl]n

lt t1! ,rtnntfl L.u LltYtt 9t1]qqr.l!611]triâ&#x201A;¬!

LLoilrfi lrroiqrdrloLilurnrnruroL6n 6r!ru'lriarntinfi Ioqio\Jrih rnlrtlo narrLLa :ruro1rlqj'lurirr:lnrtrqo riu . ! . i t 1 t R 1 1 a1' t: n 0 g l 0 0 L u u ' l l q o t | n . !

ua: rGrlor rihr uro"hqj6o:r aariui{ Lrn:rr.r'hi"hqqr.Juro.:fl do'Lrlr4r"lfr irluanrvrm:rrd aru:nnronr:rii'Ifi

;@irrc,d,;ti

\-1-l

,qq,

rrnn:d-rxr'rrnA!ri!i$d:xi j\d'ldnrd! ::u:t-lat o il

drr#r6od'r'l4o16oouror"rdr'mrrrLm inerr Lndartrrrier Ltas\iinr:oror vn-rlr ntfrr or"r otir.rtLdrt.ur6r g Llr1fr.ir,,rl 11aran LLdo.rufrdlfrn11 zuo.rdoiuh'lo r3o 1 .'ooqqr.luur6ou6l nri'rrrfrnurdargrrr'nur,uolnrn rir drtYurrir u6l rraroonririuuo.r ft6lJUlnt:nuflrs(o.i 1n0nF:.i .u0 n oahsru:R'rs!onrolfi1aro r (rlairud: grrLruraLnoilo mnn:qir.rniv rirdllunrnoonlnil iLndf, nsrn6ir:'ir rilour:orir:r6in da.rndo.roar::ariao.r z z , ulR t')n1nn1n:!Ltnit!ll-t -ni1.lft1.l ronlrniti.riorvro drgRrr'{u! ru 0 t r t O 1 1 . 1 1 ' l . l F l X l t l l t l S l . i l tdqol - o o o , . t r "hrirrrdrurrrarr6lrra:narl6u LrJoiL4u6' Ltail:'lriarur:odr:.roqi'ld0taD:dlnql ufinur l00riy|1D0-r rur ol1 nu1l:a unflLllrllnnanlr

t!0.in1R1un:q0itil!'to-t't!M1t01ttl

nIt!gtlu

l t IJu! 0n n d tffllll t1tll

flltlfldolJlt9l-lu

tlloin1fl tt']1lflq10.t -, "4-"

lunr:rta"h nrn"lnt-Lilvnrrhra-:tncr J

n1vtll-lnu0alv oangltn6l .un nM1l

oo LUolt.ruo

{--,d-.-.ib

aRl tt2oootTalnt)(J.!n0nllollts')01 A

nt ntM1tfllntl

'hitrird'u e:fia{d'{LnnLihfi rqu Y" et t ,-l]1lntl5nfil1100nft t9rooo)n1:

irrlltdartrnrfi onrnFtnrirnir"

nYl,!l]tt tlt1i5nfl]t\llnta l 1 alu:Rnll

r:wqrlr!1nrfl6.inii r^rar6nrrrq.r

nrnordooqj rrr-n\rloo:liu raohrir.r

LUOOU tU6nTr:Lrtdn1ll

i-4!. !

f l t L r o o i i o L r dr L L o o . l i ] : nr r i : i i u

]ia,

t111LV^UO0l{Ini I (lll9toYi.t

[fi ?06!]o "a_nsfl:rri!ri'! fi i1o.r v

tlldlltfl Lndd!11t1]dnt []oa0!YltLlN

t11fl14LLflnO1:nrnlld:

!rR.l tuuounnidr!tdgntt!11.J mogtI

oova.l

-z

R15141.31U10.10',t!',ti!n1o t1

"bJhunnueio:mudor.rdroqjor:r"lon :roGr Lra:r:lrirfi oanr-::qt!t!rnre rirsingrFrrr|.rrt!{ir fir"lfinrnotirrun lldllqt I 1 1Dt lU tUtU tutli:nf|ll,lltolt 4 -1]1{U1Ull.lfln95fl !tqt',trYUl]0.tO',trU -

h ,

td 1Ll l9rt lr I11100I LLn: Lt.l1Fra 011111 )

run::y"rornraGrtdlnrrSofirllrqrn

u.J

1lt tllRl:flotRlg m}]unn nl:Lnna!11

,uorrrnl:'lrinrior rrndrtinriur:r6u ! -h l J L 1 l 6 : J u 1( | n : L O L f t l : n f l H r I

urlrd r

rRt] na:n0n1 1lrrnuo)0oRr.fr',!0 "rfiot:rlou r Rrnr: [fix o.!1.li! .A on50t14ir0 oonnr-r uh 1 r'nl:'hi:l: varnnihLEo {,

an!

'rqsodr.ifl fiidr!ro1,1r:lurrn'1flrr

-A

dlritalu anitai

u3,'t*nuuli.,i',

r:.u:v L:rLri!nnorrvrntilulq?L{rt

tvadR nn!tntgDlau

aflfltmllllutlLtdi{ullogtfllo

| t a g n 5 ! 1 at a a 1 l t v l n o n a l g

tiorrrnriltnryLtrillrn'lfi t:taol1{ ars16!a:fr onri:oueio::l1ir.ift r f.! !"

n]l tt]1lno10.t 1nt 0 talu:-g1j11

Eiui!rnrauj!e oFn:rqooonnrn

,.1 L r ! r o 9 ) , 1R l ] d n t o r L t o _ R r t L l l d 0 u I d.i1o.in11:.rrilrId.3"o-rfindrroou tr,,

"-:

"or"'lldriornrrrn6ou1.|irrorni6o rir 1 ro:firrnniraororr6rtr

, :,!"-z l 0 . r L a n . l 1 1 f t , , l 1 l . t t 1 l 1 1 . i 6 t f i 1 1 1 . t o - firrtr".rotir.r.irooro" DJ

. i-.l ntnl1]lt:ln'tnnRu ta0o ltnR1,l1111.1

(lttgtt{ta:tltl { 3i!{dfi o!t:r! .ra1l::11111111r19lnf fl 111lr114n',i:nr

nn0 r l fl n:t]ju1{11,1: ll1ttn fl1a1 6oqiani!yri L6oo"l6'y3olrilosLoyrr:

l0. tat1lanuJ00nMn1no't1l1t.!41fl 1

n'rn vs Ru

Irnoirorudorludodrrrood "nr:finur'luqir.rri::tna nrn ndu :rngirr"nrdr6'.r rdrmo'bJ'l'1mr5arrrrorilurrr:rirl:olrlniorluir vJr tu, ratSnt )fl llt5ff" l,\11lllllt0tntlt

dt l1d tatfl I t4ttfi zu)1.tu:l]nlt!11l111

,!"

atiLaro

F n I U t O L O n L l 0| l O L ? d - r 1 r U a t l : J l r F

fln!

Sioroti'luo"rvrnururfr araa.u6aur,m".r rrnoor6ooIoiti{'lriin]:dnsrrisr nirnr:Lilumr:irirl:ntrnu"otir.r'l:

nlqnlJu flronr:u llr t unnunrl L1rafr nnrr *unrFidLnnroano nl lrr4.ilrutnRnRntglll!n.!a'tlutuu

ttll

t 1 t a R t l l l 1 6 L. !u u o : I u 8 1 n :

r.irdr11!el-ro!n!Hij

nD]lln?!l]111fl LOnaoolt 611-rrlrL1ro1rll

qorl td:qnufiiaalinirrrjr rfronrrrr rirllr"ld"l d'nfirrnrr rfiovrrrn !, g l:ttlutnltn"h orortlfiotor6r.rt6 {l.J0.ti1f]loao11Jlt]111 Y1UfO l OOtA00

Rntu dl1.t5llo:L{U tn0-lttn I tl] tuu Llito

rixqqdr uo1lrqqufi.r ornmLrii.r n: lrlnuttnrnu lnuqoLLd.r nrnr:orru

ulniordrarrr:ntirtrnooardnu"ld

f)tlfltllltnt

a: d R! fll3 Lolt u dll Lt) 1.1

d t lt 141fldl U t t fl L U1l111:Uit r LliULrFr , i i ue- { 3 tL0ft1lllLfl u.imnt to:llt$01l1.lftl1a

lll0Utuli:L)lllt111l I1IUOUllLRanln_lt

0u1l4.i11110t1.1f!t01ntR11!11 1l.!nT ,. J,, ! l ) a l n L n : J L r r u o f lu n : L ! L n r : l | u L r 4 fal r

Llrlllltll.rlolrLr rnn:'l:arafi

,nfi*n',orir1i lreinifn?d'ntlt,tntri.iL6o:61"LriqoiuL6oon3.r 1 nnrrl

t{-,e,"e,

l9rou1r5d

f.t

t1{:l:nfl!

:flItn!.u0i

tnn:n1]n:.t

tL6r'hloon16uoio

fi:Jnt L9lfllttoin!n I g 4"

Yd-jnrnrfi uerioul.ido.iqrnd! !lnlta

ntYat'tllli ltntotioltl]loltt0ll

t.ut[f]u

.rinr:6ronoia:loirl nr1*!rr6'rvfr 1 flai

,t {4 1n ryir oLnaflfi

fl L:J:]n:Lrlu

|l lg )fiLtFlt1!vatnva1uo11.l1uoI't1.1i

0 ' l u ' l i L ? I F I: " u ] l o 1 u a 1 l 0 . i 0 l l m n t i

ifla:11r1rlull:i:r fi llrfr arl111rllrfi0l6n

d o 1 1 , , { ) ntut t . l i t L ! 0 I u t v l u l ! u u t t d t l ] tl|111J

o0l

LLltoLl(llu lsllotou)Ll1t1lll

tl]t

rrirriur|o:fi\ririoonir tstio

uoR

xrayigni:nruod"lrlr.ufl nrnr io o'dril

t.]n.rnl1]ll],Tr5fln:r] flntl11]1n

tLnil]'t.l11lo1!!n:!nt !R')tn1-11.i tn0]l n f l r F r : l d . i l n t r t ! 1 i t l u ut u t : 1 t l

ruoo!-il "uisarriao-eo LrloiLdrd

n,r6fiun'ilrr-r,ra,rn,oin,n,r LLriLroirrn Lu"ln{rlrlnrrfi drrlrlr

uoRnlni L"ll]u".iylxfiad"{ In0i r

r zu:fr sr"r ooirrrrdii'lniiioodr.rrla-r o : a n u n R ! l t l a ' l n L o v l t a n ! t u i ! t l 'lriidaiutatrLLpinrnooL6ooLdu'Ll6rtr flfl1r u1nulirrll1rl,1Ifr1.rolllrlfl11Lo Ll -l

ddu TtR t!tl]o t Lru L! l9rttt9tdut9lta a l u o r L 1 l l 1 r u1 | I B 1 r O i . ! n t . t -i uL f n o . !

-Y:'

I { .! d 1 0 l F r o L U L l ] t d l ]L n f i R ! t ! 1 ] 9 0 L U

n d u d ot d n a ! 1 ] s i n o t n t i l t n ' 1 1L 1 l t d n

-illraullrirnrrflnurqioll .t,

aruorfidra tfiodi':oniui'Libl to rio ,tra1i d8t lfl Lnu ttt||lJ)

rfltfl110ullu

allnoLr.rlr0rnull0tdllntrlr,irlnnrrnfr!1

8 r_tulrunt:l,rrrrriou"lui:rLna"lr L "[ ru:l]dr:.] r )-flr1lo Lfi tJlo Ll^lu0l l i,ll r

fl lndna (genus)1 ri.irr.1:-ririr:111fl

arirarrunrnln':rnr,rrla,urarr' nrrrYLLn : Ffl!ruI!!runl'ihi,r'rirrlf r6rnrnl! lrrrtrntru (BRT)

t1luotuo0 I tetnhdel|rntRLolrfl 1,{ridnaHacmadipsatfi uoriiL6tlr .-l

qrwrrLLriru6Lu1ttt! arn'jsr6r'inurna:inornrani

toullllttulto0tto:rn0tl]ltfntn1] ]ltltu

f l a l n t n L d :n t 5 0 l f l t n t L d : r . u u t f l n I I

adru{rJ6.rfirrd.rhti i filrurirh -

!:,:

duLn Llrnar,)nrarJnu irn:ru'rr flr a

-t

{

" fin!tnu r?:|dotnq!! trrnttaird r )

n1i rnal'na ftt1ta.,vfra n1t u1'ti nt' i1 u G t tra"u ai ti{d n i r m dnnr; a t t r: x ]n fl 1nIt1 1d H tm

1dtp\ tdle n fl L

'lu1lr: LndIna \aL 61{lt ilrJdr:rdi!).ri

r"rLnia n"lnri--rrrr:"h tfror'hod.r'hitn o nru:inurmaoi rms.rn:nirn:nur;u -.'i rnn ollltnRonllnollmttYuonla.xt

uuii,,r,-ouuui,uu.rr,i,o-q1,J,o,r v, utll d0)na 50nt:nLl'{!n:-11

' ' \ r n " A d v a . n c cl h a i 1 . n d Ceosnpi;. lou iir: dirlnuG iiunr!.in dlrru 6rr;i!l qarrr! L'lir drulxrtl